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Lab 2 + 3 background info
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What are specialized receptors?
Receptors in the muscle that respond to the stretching of the tendon attached to the muscle and then send signals to motor neurons through a single synapse.
What happens to the muscles in response?
They depolarize and twitch (contract) in response to the incoming impulse from the motor neuron.
What do the skeletal muscle receptors convey?
Information about muscle length, tension, and pressure to the central nervous system.
What are sensory receptors responsible for?
Providing information about the length or the rate of change of the length.
What muscles do sensory receptors convey information of?
Muscle spindles
How are muscle spindles arranged?
In parallel with muscle fibers.
How are the muscles spindled stretched?
Stretched when the muscle is stretched by an external force.
What is the purpose of muscle spindles?
Develops antigravity reflexes and maintains muscle tone.
What do muscle spindles contain?
Contain a small bundle of intrafusal fibers and extrafusal fibers
Do intrafusal fibers contribute to the overall tension of the muscle?
No
What do intrafusal fibers contribute to?
They regulate the excitability of the sensory afferent spindle nerves
How do intrafusal fibers regulate the excitability of afferent spindle fibers?
They mechanically deform the receptors
What are intrafusal fibers innervated by?
Gamma motor neurons.
What do most muscles consist of?
Extrafusal fibers
What are extrafusal fibers innervated by?
Alpha motor neurons
What are extrafusal fibers responsible for?
Developing muscle tension
What happens when a muscles is stretched?
The excitation of its muscle spindles cause a reflex contraction of the muscle
What is this reflex called?
Stretch (myotactic) reflex
What causes minimal delay between the muscle stretching and the reflex contraction?
The monosynaptic pathway.
What happens during the monosynaptic pathway?
Afferent nerves from the spindles synapse directly with motor neurons
Are there interneurons in this monosynaptic pathway?
There are no interneurons
What does this monosynaptic pathway constitute to?
The shortest possible reflex arc.
Why do we have this reflex arc?
Its the main reason why we keep our balance and do not fall when changing certain body positions.
Which muscle groups are involved in plantar flexion of the ankle?
Gastrocnemius and soleus
Which muscle groups are involved in dorsiflexion of the ankle?
The anterior compartment of the lower leg (so the tibia’s, extensor digitorum and hallucis longus, and the peroneus tertius.
Besides excitatory inputs from stretch receptors, what synaptic inputs might influence the activity of spinal motor neurons?
Descending motor commands
Inhibitory spinal interneurons
Sensory afferent neurons
Modulatory inputs
How many muscles does the human eye have?
6 muscles
What surface are these muscles attached to?
The exterior surface
How many groups are the muscles in
3 antagonistic pairs
How are these muscles grouped?
Horizontal
Vertical
Torsional
movement and position of the eye
What muscles move around the vertical axis?
Medial rectus and lateral rectus muscle
How do these muscles move around the vertical axis?
Medial rectus: Turns eye towards nose (ADDUCTS)
Lateral rectus: Turns eye away from nose (ABDUCTS)
What muscles move around the horizontal axis?
Superior and inferior rectus muscles
How do these muscles move around the horizontal axis?
Superior rectus: Turns eye up w slight rotation to nose (ELEVATES + INTORTS)
Inferior rectus: Turns eye down w slight rotation away from nose (DEPRESSES + EXTORTS)
What muscles move around the torsional axis?
Superior and inferior oblique muscles
How do these muscles move around the torsional axis?
Superior oblique: Rotates top of eye to nose w slight depression (INTORTS)
Inferior oblique: Rotates top of eye away from nose w slight elevation (EXTORTS)
How are these muscles innervated?
Innervated by motor neurons that have electrical activity w a tonic and phasic component
What does the tonic component control?
The position of the eye
What does the phasic component control?
The velocity of the eye
How are the eye velocity commands sent?
Along direct path from specialized brain formations or fields to the motor neurons
What are the eye position commands a product of?
The integration of eye velocity commands sent along an indirect path to a network of neurons functioning as neural integrators.
What is the output oof the integrator?
Provides eye position commands to the motor neurons
What do the integration of signals control?
5 types of eye movement each w a unique function and distinctive property
What are the five types of eye movement?
Saccades
Pursuit
Vestibular ocular reflex (VOR)
Vergence
Optokinetic reflex
What is the recording of the electrical activity of the eye known as?
Electrooculogram (EOG)
What is the fovea centralis (focal point)
The region of the retina that sees in detail.
What do the saccadic eye movements do?
Rotate both eyes of that the image of interest falls on the fovea.
How fast are saccadic eye movements?
They’re quick w a velocity as high as 800 degrees in a second
Are the saccades accurate?
Yes
How are the saccades accurate?
The system uses an internal estimate of eye position from its neural integrator to guide and stop the saccades.
What does the pursuit eye movement do?
Keeps the fovea pointed at a moving target like a bouncing ball
What happens during pursuit movement?
There is an initial delay because the signal from the eye indicates that the object is moving, which has to be conducted through many synapses to the brainstem.
How does the saccade help with pursuit movement?
Helps the fovea catch up to the object until the pursuit movement begins to track the object.
What does the VOR movement do?
Keeps the image of the outside world stationary on the entire retina when the head moves.
Is VOR tonic or phasic?
Its a phasic response that is faster than pursuit
Why is VOR faster than pursuit?
Because it’s a simple central reflex arc that involves only three neurons.
Where does the signal that indicated the velocity of head movement originate from?
In the semicircular canals of the ear which goes through an afferent nerve and an interneuron on its way to the motor neuron of the oculomotor muscles.
What happens during VOR movement?
The muscles rotate the eyes at a velocity that matches the velocity of the head, keeping the image stationary on the retina.
Are the eyes held on the image going through a tonic or phasic response?
Tonic response
Where is the tonic response happening?
The tonic response is along an indirect path through a reverberating neural circuit between the afferent nerve and motor neurons
What would happen without this neural circuit (short-term memory device)
The eyes would drift back to the centre and off the image while the head was still rotated.
What is the vergence movement?
Points the fovea of each eye onto an object when you look from a target that is distant to the one that is closer (vice versa)
What happens during vergence?
The eyes rotate in opposite directions
What happens to the eye when looking from far to near?
Convergence
What happens to the eye when looking from near to far?
Divergence
Why is vergence important?
Prevents double vision
When is the optokinetic reflex activated?
When the full field of vision has moved across a large portion of the retina
How fast is the optokinetic reflex?
It is slower than the VOR, hence complementing this movement
Why do we have the optokinetic reflex?
Better suited for working on slow and prolonged movements
What device is used to test the optokinetic reflex?
An optokinetic drum
What does the optokinetic drum do?
Projects a series of rotating images on the wall of a small circular room while the subject is seated in the centre of the room/