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mechanism through which one organism alters the environment to impact another organism
biotic interactions
what are the two type of interferences that create environmental change?
removal and addition
when one organism removes something from the environment or reduces its availability
removal interference
what are the effects of removal interference?
usually negative for one individual
what forms of interaction are considered removal interreferences?
competition, parasitism, herbivory
use or defense of a shared resources, reduces availability, two species occupying similar niches (-,-)
competition
do farmers assume competition in agriculture and if so, how do they deal with it?
yes, they flood the environment with the resources
organisms living in close proximity where parasite derives nourishment from the host but does not kill it (-,+)
parasitism
when plant tissues is removed from the ecosystem (+,±)
herbivory
negative impacts of herbivory
less biomass, less photosynthesis, nutrient removal, decrease in yield
positive impacts of herbivory
stimulate new biomass growth, change species composition
strategic movement of livestock among partitioned pastures
rotational grazing
benefits of rotational grazing (4)
reduced weeds, enhance forage and animal productivity, better nutrient distribution, less compaction/erosion
when are organism adds to the environment and impacts other organisms
addition interference
organism living on the body of another organism, one organism provides habitat for another, no nutritional benefit, form of commensalism
epiphytism
examples of epiphytism talked about in class
algae, moss and ferns, bromeliads and vanilla orchids
when both organisms make additions and both benefit (+,+)
mutualism
degree of specificity of mutualism (2)
facultative and obligate
facultative mutualism
take it or leave it
obligate mutualism
one on one necessary
examples of mutualism talked about in class (3)
pollinators and plants they visit, legume plants and Rhizobia bacteria, plant and mycorrhizae
production of a compound by a plant with an inhibitory or stimulatory effect
allelopathy
combination of a species requirement for every physical and biological resource, includes habitat, nutrition and relationships
ecological niche
most limiting requirement determines growth rate or even presence of the organism
Law of the Minimum
niche requirements overlap, species compete and restrict one another
Competitive Exclusion Principle
all possible conditions in which a species can exist
potential (fundamental) niche
niche that the organism actually occupies due to resource acquisition
actual (realized) niche
_____ partitioning allows organisms to divide up a limiting resources
niche
evolution of a trait in response to competition for a resource
character displacement
change in the trait exhibited in the absence of competition/selection
character release
how are niches used in agriculture?
to determine what plants will grow where
we should intercrop plants with _____ niche requirements
different
competition between different species
interspecific
competition among the same species
intraspecific
what are the positives to decreasing the space between plant rows?
increase plant populations and yield, improved competition vs. weeds
what are the negatives to decreasing the space between plant rows?
difficult passage for equipment, possible increase in plant diseases
ways plant compounds can be released with allelopathy (4)
volatilization, exudation from roots, decomposition, leaching from leaves
allelopathic compounds (4)
tannins, phenolics, terpenes, alkaloids
what are the characteristics of compounds for allelopathy (4)
compounds short lives, may interact with other compounds in environment, differential effect on plants, concentrations vary with plant and conditions
what are the steps to demonstrating allelopathy is occurring? (6)
show compound is released from plant, compound reaches toxic level in environment, uptake by target plant, identify compound and physiological basis for response, understand interaction with environment
what is the negative impact of allelopathy called and what does it do?
inhibitory effect, phytotoxins suppress plant growth
aside from allelopathic plants competing, they can also decrease _____
germination, emergence, growth, soil, symbionts
what are examples of inhibitory allelopathy we talked about in class?
bitter grass with corn, lambs quarter and bean crops
how is allelopathy used for the biological control of weeds? (5)
crop itself, mulch, cover crop, intercropping, crop rotation
what is the positive impact of allelopathy called and what does it do?
stimulatory effect, increase plant growth by pairing with other compounds
what are some examples of stimulatory allelopathy that we talked about in class?
corn cockle and wheat, alfalfa on various crops
predictable changes in species structure and community composition over time
ecological succession
trajectory of succession altered or restarted by ______
disturbance
3 characteristics of disturbance
intensity, frequency, scale
what are some examples of disturbance?
wind, water, temperature, disease epidemics, any agent of change
what agricultural practices are considered disturbances?
tilling (cultivation), planting, harvest, burning
what characterizes the early stages of succession on an agricultural plot (annual weeds)?
rich sources of nutrients, light, harsh and unprotected, not diverse, no vertical structure, early colonizers change environment, life cycle completed in single season, recruitment from the seed bank
what characterizes the first 5 years of succession on an agricultural plot (perennials established)?
multi-year lifecycle, good competitors, out-compete annuals
what characterizes the first 10-15 years of succession on an agricultural plot (pine colonization)?
tree seedlings sprout, shade tolerant species
what characterizes the first 50 years of succession on an agricultural plot?
mature pine forest, hardwood colonization
what characterizes the 100-150 years of succession on an agricultural plot?
oak-hickory forest
where does primary succession occur?
colonization of bare rock, lava flows, glacial retreats, dunes
where does secondary succession occur?
occurs on pre-existing soil following disturbance
what are the characteristics considered for successional species identification? (5)
life history, colonization, dispersal, reproduction, competition
characteristics of colonizers (6)
adapt to extreme environments, wind dispersal, seed bank, low diversity, poor competitors, fast growing, reproduce quickly, semelparous
characteristics of persisters
more restrictive habitat requirements, animals dispersal, high diversity, good competitors, slow growing, reproduce slowly, iteroparous
2 factors that initiate succession?
allogenic and autogenic
what are allogenic factors?
external
what are some examples of allogenic factors?
fire, climate, weather, grazing, human-mediated changes
what are autogenic factors?
caused by organisms in the community
what are some examples of autogenic factors?
facilitation and inhibition
diversity and productivity of natural systems is greatest when moderate disturbance occurs periodically, patchy landscape with multiple successional stages
Intermediate Disturbance Hypothesis
how does modern agriculture deal with succession?
ignores successional process, utilize inputs to replace harvest, rely on early successional stages with high net productivity
how to improve sustainability with succession and agriculture?
less reliance in inputs, utilize later stages of succession
plant or retain trees in crop system
agroforestry
what are the 4 ways populations change size?
birth, death, immigration, emigration
population growth in an unlimited environment where growth continues indefinitely
exponential growth
is the exponential growth model density independent or dependent?
independent
what are the 5 assumptions for an exponential growth population model?
unlimited favorable environment, closed population, continuous growth, no age structure, constant birth and death rates
realistic population growth marked by a carrying capacity and max amount of population an area can hold
logistic growth
is the logistic growth model density independent or dependent?
dependent
carrying capacity
upper limit to population size that an environment will support
if P is very small (near 0) then the growth rate of the population is…
1 or exponential
if P is very large (near K) then the growth rate of the population is…
about 0
what are the added assumptions for the logistic growth model?
constant environment and constant carrying capacity
what happens as a population approaches K?
fluctuates due to time lags
can K change?
yes
how do we impact the carrying capacity in agriculture?
modify environment so crops can increase beyond K
what is the life history strategy used by the following species: colonizer, devotes more energy to reproduction, high rate of increase, responds to disturbance, colonizes favorable situations quickly
r strategists
what is the life history strategy used by the following species: persister, devotes more energy to maintenance, longer life spans in more stable habitats, lower reproductive rates
K strategist
sequence of events in decomposition (4)
organic matter (litter), detritus, humus, inorganic compounds
what directly decomposes plant matter?
bacteria, fungi, actinomycetes, termites
what directly decomposes animal organic matter?
insects and other organisms
what are the 3 domains?
eubacteria, eukaryotes, archaea
what do soil bacteria do?
break down plant OM, mineralize, nitrify, denitrify
what do soil actinomycetes do?
filamentous, mineralize plant OM
what do soil fungi do?
mineralize plant OM, enzymes break down lignin
what dose soil mycorrhizae do?
forms mutualistic relationship with plant roots where fungi provides P and plants provide sugar
what do soil metazoa do?
break down OM into smaller particles, some mineralization
what do soil earthworms do?
eat soil, feed on bacteria and fungi, improve soil structure, increase microbial activity in soil
what do soil termites do?
breakdown wood with high C:N ratio, mineralize plant OM
what do soil millipedes and pill bugs do?
feed on organic matter
what do soil mites do?
fungivores and predators , very diverse
what do soil collembola (arthropod) do?
fungivores with and abdominal appendage
what do enchytraeid worms do?
eat fungi