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clade
mrca + all descendants
monophyly
mrca + all descendants
paraphyly
mrca + some descendants
polyphyly
no mrca + some descendants
synapomorphy
a trait that all species in a clade share, which evolved on a branch leading to that clade
homoplasy
similar trait not due to ancestry
2 primary sources of macroevolutionary insights
1) paleontology
2) phylogenetics
biological species concept
species are a group of interbreeding natural populations that are reproductively isolated from others such groups
does reproductive isolation have to be absolute for the BSC
no
allopatric vs sympatric isolation
speciation occurs due to geographic isolation vs. in the same location due to mutations and reproductive preferences
processes where reproductive isolation may occur
finding mate —> fertilization —> development and growth of zygote —> adult survival and reproduction—> growth and survival of offspring
define prezygotic barriers
barriers that prevent successful mating and/or fertilization
types of prezygotic barriers
geographical, ecological, temporal, behavioral, mechanical, cellular
how are apple maggot flies an example of prezygotic barriers
some flies started preferring introduced apples over native hawthorns —> temporal isolation led to divergence
types of intrinsic post zygotic barriers
inviability, sterility, abnormal development of zygote
extrinsic post zygotic barriers occur when
hybrids are less fit in an environment compared to either of its parental lineages
ecological speciation
speciation because of adaptation
adaptive radiation
process of adaptations generating diversity
4 features of adaptive radiation
1) recent common ancestor
2) phenotype-environment correlation
3) trait utility
4) rapid speciation
adaptive radiation is caused by
1) ecological opportunity (e.g colonization, extinction of competitors, innovation)
2) high propensity for speciation
ecology
study of how organisms interact with each other and their environment + distribution/abundance of species + structure/function of ecosystems
population
all individuals of a species in one place at one time
community
all species living together in one place at one time
ecosystem
all species + their non living environment
factors that determine species distribution (biggest to smallest filter)
dispersal, abiotic conditions, species interactions
lethal zone
portion of environmental gradient where species performance equals zero
ecological niche
combination of physiological tolerances and resource requirements of species
hutchinsonian niche
an ‘n-dimensional' hyper-volume in which each axis is an ecological factor important to the species
temperature is mostly a function of:
latitude
what factors determine biomes
temperature, rainfall, and the seasonality of them
factors that influence rainfall
atmospheric circulation, currents, rainshadows
cell order (0-30, 30-60, 60-90)
hadley, ferrel, polar
intertropical convergence zone
the zone of rising air near the equator that creates the hadley cells
why does the itcz move more over asia
more land mass
doldrums and horse latitudes
areas of low wind at the equator and 30 degrees
roaring 40s
strong westerlies in S hemisphere due to low land mass
deserts are near
cold water currents
critical max/min temp
temp where some critical function of an organism is lost
fitness is maximized when reactions occur at
optimum temp/osmotic conditions
poikilotherms
can’t regulate internal body temp
homeotherms
can regulate internal temperature
bergmanns rule
homeotherms are larger at higher latitudes
allens rule
homeotherms have smaller appendages at higher latitudes
enhanced vascularization purpose
redistributes heat, useful for cooling (loss thru convection)
countercurrent circulation
helps conserve heat in cold environment organisms
whittakers diagram
high precip and high temp = high productivity
hypothesis about complexity
greater complexity arises form greater cooperation among previously independent units
selection acts at a specific level when there is:
variation across the level, units exhibit heritability, units have differential fitness
mitosis benefit for cooperation
reduces variation between units
meiosis benefit for cooperation
randomness prevents any parent lineage from outcompeting the other
single cell beginning benefit for cooperation
less competition initially
meiotic drive
biological process where one allele is biased in transmission because the biased allele prevents proper formation of the other allele
if a favored allele is bad at the individual level, an organism may:
evolve silencing “restorer” alleles
transposable elements
self replicating segments of DNA, separate from cell replicatio
how do genoomes not explode from TEs?
alleles arise that methylate DNA, balance between high level and lower level selection
3 methods that cells stay cooperative
1) starting from single cell
2) separation of germ cells + limited divisions in germ cells
3) tumor supressors
weed resistance can evolve through these 3 ways
1) pre-existing variation 2) de-novo mutations 3) gene flow
3 axis of adaptation probability model
mutation supply rate, wild type fitness, probability of adaptation
evolutionary toggling
once resistance evolves, stop taking drugs so that non-resistant strain takes over (cuz fitness cost), then reapply
evolutionary rescue
the process where a population facing extinction due to environmental stress is saved by adaptive evolution
probability of evolutionary rescue depends on
population size, how much fitness was reduced, beneficial mutation rate
what mostly influences seasonality at high vs. low latitudes?
temperature vs. rainfall