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Who introduced the payoff matrix?
Hamilton
What is the payoff matrix, and how it does it illustrate the cooperation problem
it categorises social interactions (actor, recipient) into mutualism (+, +), selfishness (+, -), altruism (-, +) and site (-, -)
Considering the payoff matrix, why should altruism “not evolve”, what is the downside of it?
the actor suffers a fitness cost while benefitting others, reducing own reproductive success
What individuals undermine cooperation
free riders
What is the free-rider problem in cooperation?
individuals who exploit cooperative benefits without contributing
Why does mutualism still face the free-rider problem, since both parties benefit?
individuals may attempt to minimize their own costs while maximizing benefits.
What is kin selection?
A mechanism where individuals help relatives to increase their inclusive fitness.
Who proposed inclusive fitness theory?
W.D. Hamilton
What is inclusive fitness?
the total fitness/genetic success of an individual’s genes
How to calculate inclusive fitness?
direct fitness (personal reproduction) + indirect fitness (helping relatives reproduce also pass on own genes)
What is Hamilton’s rule?
the analysis for altruistic behaviour as an act has a cost for the actor and benefits for the recipients
When does altruism evolve, according to Hamilton’s rule (equation)
c < r x b, where c = cost to actor, r = relatedness, b = benefit to recipient.
What does r represent in Hamilton’s rule?
the probability two individuals share an allele by descent
What is the “green beard effect” (though it is rare)?
Genes produce a trait, recognition of that trait, and altruism toward bearers of the trait.
What is the genetic relatedness between siblings?
Approximately r = 0.5.
Why is kin selection insufficient for explaining cooperation in large human societies?
Because many interactions occur between unrelated individuals (kind to nonkin).
Functional significance of kin selection in cooperation?
Promotes altruism toward relatives
According to natural selection, who are individuals expected to behave kindly towards
relatives, promote genetic selfishness
7 reasons why we are kind to nonkin
reciprocal altruism, mutualism, indirect reciprocity, generalised reciprocity, negative reciprocity, strong reciprocity, cultural group selection
What is reciprocal altruism, according to Trivers
direct form of reciprocity, individuals help others with the expectation that the favour will be returned in the future
What are 3 conditions required for reciprocal altruism?
Frequent interactions, ability to recognise individuals, and contingent behaviour.
What is mutualism?
Immediate benefits to both individuals without delay or reciprocity.
Why might mutualism be mistaken for altruism?
Because both parties benefit, but motives may be selfish.
What is indirect reciprocity?
expectation that the favour will be returned, not necessarily by the recipient but by someone else in the community
Two types of indirect reciprocity
downstream and upstream
What is downstream indirect reciprocity
helping others to build a reputation
What is upstream indirect reciprocity
helping others as a form of niceness
What is generalised reciprocity?
Altruism without expectation of return
What is negative reciprocity?
Punishment for harm done
What purpose does negative reciprocity fulfil
deters free-riders
What is strong reciprocity (two actions)?
Cooperate with others and punish non-cooperators
What is the benefit of strong reciprocity
helps to maintain social norms and cooperation within a group
What societies rely heavily on reciprocity?
Hunter-gatherers like !Kung San, Aché, Yanomamö.
What evolutionary human behaviour is reciprocity essential for
fair sharing of resources
What does the cultural group selection principle propose
that groups with altruistic tendencies will outperform less cooperative groups
What 3 cognitive demands does reciprocity impose?
Memory, recognition, and contingent decision-making.
What is game theory?
Mathematical framework for strategic decision-making in social interactions.
What is strategic decision-making
outcome for each participant depends on the choices of all involved
What are 2 processes involved in strategic decision making
rationality and utility
What is the assumption of rationality in game theory
that all players are rational and they know that all other players are rational
What is the assumption of utility in game theory
utility maximisation where players will consume a good to satisfy a desire
What is Nash equilibrium
a set of strategies where no player benefits from changing their strategy while the other players keep theirs unchanged
What is an Evolutionarily Stable Strategy (ESS)?
A strategy adopted by a population that is resistant to invasion by alternative strategies
What is the prisoner’s dilemma?
Game where defection yields greater personal short-term gain but mutual cooperation is the best collective outcome
What are the 4 outcomes (what is the acronym, when you fall?)?
TRPS
What is T in TRPS
temptation to defect (0 years, 20 years)
What is R in TRPS
reward for mutual cooperation (1 year, 1 year)
What is P in TRPS
punishment for mutual defection (5 years, 5 years)
What is S in TRPS
sucker’s payoff (20 years, 0 year)
What is the dilemma that involves a repeated game of game theory, allowing strategies to develop based on past interactions
IPD
What is IPD
Iterated Prisoner’s Dilemma
What is tit-for-tat (TFT) strategy?
Cooperate first, then copy opponent’s previous move; kind, retaliatory, forgiving.
What type of strategy is TFT
ESS
How does game theory apply to real-world cooperation problems?
E.g., climate change, resource sharing.
Functional significance of game theory?
Explains strategic behaviour and evolution of cooperation.
What is “cold cognition” in cooperation (+ 3 examples) ?
Reason-based processes
What is “hot cognition” in cooperation?
Emotion-driven processes influencing social decisions.
What role do emotions play in cooperation?
Serve as heuristics and “thermostats” to guide social behaviour

What is this table demonstrating
fortunes of others emotions
Positive feelings + positive feelings = ?
symhedonia
Positive feelings + negative feelings = ?
schadenfreude
Negative feelings + positive feelings = ?
jealousy
Negative feelings + negative feelings = ?
empathy
What is schadenfreude?
Positive feelings at others’ misfortune; can undermine cooperation.
What is symhedonia?
Positive feelings at others’ success; promotes prosociality.
What two feelings, according to the fortunes of others emotions, promote pro-social behaviour
empathy and symhedonia
What is the tragedy of the commons?
Overuse of shared resources leads to collective ruin
What is a public goods game?
An N-player prisoner’s dilemma testing cooperation in resource contribution.