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Evolution
change in frequency of a trait through time (trait = genotype of phenotype)w
Microevolution
change in allele frequencies within a population over short time scales
Macroevolution
large-scale evolutionary changes above the species level resulting form long-term microevolution
What are the three types of evolution paces?
stasis: constant linear
gradual change: increasing linear line
Punctuated change: straight line —> rapid increase —> straight line
What are the mechanisms of evolution?
Mutation
Gene Flow
Genetic Drift
Natural Selection
Which ones are neutral evolution?
Mutation
Gene Flow
Genetic Drift
Darwin’s Postulates
The following 3 postulates make evolution by means of natural selection inevitable:
struggle for existence
variation in features related to survival and reproduction
variation is passed from generation to generation
What are the three types of selection?
Disruptive: climate changes disrupt selection pressures on populations
Stabilizing Selection: maintain status quo as long as the environment is constant
Directional: one trait favored to another trait
Artifical: Domesticate and selective breeding
Sexual Selection
Variance in mating success of members of the same sex and selects for traits that increase the relative fitness of an individual by allowing them to secure greater mating success.
Intrasexual Selection
Male competition where dominant males gain access to females
Intersexual Selection
Femalce choice where female select most attractive male
Monogamy
Individuals mate exclusively with each other
Polygamy
individuals mate with more than one partner
Polygny
males mate with more than one female
Polyandry
females mate with more than one male
Promiscuity
individuals mate indiscriminately with multiple partners
What are the female preferences for male behavior?
grooming
access to resources
protection
Theory of Evolution
Overproduction: species tend to reproduce more than one can survive to maturity
Variation: individuals of a population have many different characteristics
Selection: individuals survive longer and reproduce more than others
Adaptation: those that survive pass on their traits to their offspring (the favored traits will be more common in the population)
Global optimum
The “best” solution for natural selection to reach/achieve in a population.
Local optimum
The “good” solution for natural selection that sometimes, (normally) reaches in a population.
What are ways humans have affected evolution of our and others species?
Habitat Destruction
Overexploitation of resources
Invasive Species
Pollution & Environmental Changes
Artificial Selection
Cultural & Tech Buffering
DNA
Deoxyribonucleic acid
Unit of inheritance that contains the ‘recipe’ for creating and differentiating cells into a whole organism
4 nucleotide bases
Adenine
Guanine
Cytosine
Thymine
How many chromosomes do Humans typically have?
22 + the sex chromosomes = 23
Mitochondiral DNA
Very short chromosome that is maternally inherited and has multiple copies
Diploid
2 copies and recieve 1 copy of each chromosome from each parent
Meiosis
2 ways that create new genetic variation
50% chance that parent will pass on either chromosome of given homologous pair
genetic recombination ensures offspring will have different genetic makeup
Gregor Mendel
Discovered laws of inheritance from breeding garden pea plants
Law of Segregation
Characteristics of organisms are determined jointly by two articles, one inherited from each parent
Law of Independent Assortment
“particles” for different traits are independently inherited. Genes do not influence each other with regards to the sorting of alleles into gametes
Law of Dominance & Uniformity
Some “particles” are dominant over other particles
Hardy Weinberg Equation
Determines if evolutionary chance is taking place or in equilibrium
p² + pq + q² = 1
Hardy Weinberg Equilibrium
Population is infinitely large
No mutation
No genetic drift
No gene flow
Natural selection is not operating
Mating is random
All members produce same number of offspring
Heritability
How much of a trait is genetic and how much is something else (environment)
H = (variation accounted for by genotype)/(total variation)
Mutation
A spontaneous change in the chemical structure of DNA
What are the three types of Mutations
Harmful: negatively affects
Neutral: no affect
Beneficial: positive affect
Gene Flow
Introduction of genetic material from one population/species to another through reproduction of migrating individuals
Genetic Drift
Random change in a gene frequencies b/c of smapling variaiton that occurs in any finite population
Bottleneck event
And even that decreases and numerically harms an original population that alters and decreases gene frequencies
Founder’s Effect
Gene flow through chance isolation
Serial Founder Effect
Explains the genetic drift of Archaic Homo sapiens in the African continent
Migration out of Africa into other continents continued as serial founder effects
Do most human variation exist within or between groups?
Within
Mutation
increases variation WITHIN a group and BETWEEN groups
Genetic Drift
decreases variation WITHIN a group, but increases variation BETWEEN groups
Genetic Flow
(usually) increases variaiton WITHIN a group, but decreases variation BETWEEN groups
Modern Evolutionary Synthesis
Evolutino is genetic and measurable
Multiple forces shape evolution
Ronald Fisher
Menedlian inheritance explains continuous traits
JBS Haldance
Mathematical equations for NS and mutations
Sewall Wright
Genetic Drifts and Fitness Landscaoe
Theodosius Dobzhansky
created definition of evolution (showed the evolution works in nature, not just models or theory)
Extended Evolutionary Synthesis
Added New drivers to evolutionary changes such as development bias, niche construction, and non-genetic ways of inheritance
Current Evolutionary Theory
Evolution is shaped by genes, development, plasticity, and environment
Organisms modify their own selection pressures
Non-genetic inheritance
Works to explain rapid adaptation and evolutionary innovation
Main characteristics of a Primate
Grasping Hands and feet
Nails instead of claws
Limb driven locomotion
Reduced sense of smell, enhanced vision
Forward-facing eyes
Large brain to body size
Long gestation
Small litters
Long juvenile period to learn
Long lifespan
Increased dependence on learning and behavioral flexibility
What are the major proximate theats to primates?
Habitat loss
Hunting & Capture
Disease
Life History Theory
Trade-offs and energy allocation
difference between fast life historu (r-selected) & slow life history (k-selected)
R selection
short lived
small body size
fast maturation
low infant investment
early reproduciton
K selection
Long lived
Large body size
reproduce at a larger age
slow maturation
high infant investment
Menopause
natural end of menstruation, usually defining the end of period cycles
‘Grandmother’ Theory
Defines how menopause is evolved by older women to increase inclusive fitness by helping raise children (increase grandmother survival by tring to have more children later in life), enhancing genetic success and explains why human females live long post-reproduction, unlike most other mammals.
Primate Socio-ecoloy
Study of how primate social systems among primate societies are influenced by the environment
Primate Dentition
(ICPM) for upper and lower Dental Formula)
I —> Incisors
C —> Canines
P —> Premolars
M —> Molars
Jarman/Bell principle
The consequences of the negative allometric relationship between body mass and nutritional requirements,
Large-bodied species need LESS energy per mass unit than small-bodied species
Reasons for Living in groups
lowers predation risk
improves predator detection
allows collective defense (mobbing behavior)
Pair-Bonding Species
mate guarding, increased parental investment,
Cooperative Breeding
Males & other individuals in group help raise the young
Infanticide
A male reproductive startegy
Predictions:
change in male residence or status and only kill insnats taht result in resumption of female crying
Data:
takeover by new male follows 85% of deaths
unweaned infants primarily targeted
Weaning
the process of energy becoming more available for the mother once nursing and dependance on mother from offspring gradually decreases
Sociality
Females baboons form strong social bonds that result in more reproductive success and live longer
Altruism
A behavior that helps another organism at a cost to the actor’s own reproductive fitness (sacrifices its own fitness for the sake of their group’s fitness)
Kin Selection
Not all members of a group are equally related and related individuals often cluster
W.D. Hamilton’s rule
rb > c
r = coefficient of relatedness between actor and recipients
b = sum of benefits to individuals affected by behavior
c = fitness cost to individual performing behavior
Kin selection
Natural selection favors altruistic behaviors that increase the reproductive success of close relatives, even at a cost to the individual’s own survival or reproduction
Inclusive Fitness
A better measure of fitness
NS favors strategies that increase inclusive fitness
(Direct fitness + Indirect Fitness) * Increased probability they share a given allele [r]