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Meristems
the region of plant growth. Primary growth occurs at the opical meristems at the tips of roots and shoots.
Primary Growth
growth of the tips of roots and shoots is divided into three zones: zone of cell division, zone of elongation, zone of differentiation
Zone of cell division
this contains the meristematic cells. These cells undergo rapid mitosis to produce new cells
Zone of elongation
the new cells no longer divide; instead they begin to expand and elongate.
Zone of differentiation
once at full size, the new cells start to specialise into specific tissues such as xylem, phloem and root hair cells
Important Photohormones
Auxins, cytokinins, ethylene, abscisic acid, gibberellins
Auxins (IAA)
produced in the shoot and root tips. Promoted elongation in the shoots but inhibits elongation in the roots. Promoted apical dominance (growth of main shoot/root).
Cytokinins
Produced in the roots. Inhibits apical dominance by promoting lateral budding.
Ethylene
Ripens fruit and promotes abscission (detachment of fruit and leaves).
Gibberellins
Breaks seed dominancy and promotes germination
Tropic responses
plant growth response that occurs directional to the stimuli. Can be positive or negative
Light
Photo
Water
Hydro
Gravity
Geo
Chemical
Chemo
Touch
Thigmo
Sun
Helio
Electricity
Electro
Tide
Rheo
Food
Tropo
Temperature
Thermo
Response Mechanism (tropic)
Tropic responses are due to uneven distribution of Auxins that cause uneven elongation of cells in areas of primary growth
Effect of gravity (Tropism)
Causes auxins to accumulate in the lower sides of the stem and roots resulting in geotropism
effect of light
causes auxins to breakdown and accumulate in shady sides of shoots resulting in phototropism
Effect of water
Auxins are water soluble and accumulate in root cells with a high water content resulting in hydrotropism
Transport of auxins
passively through the phloem or actively from cell to cell. Active transport allows for chemotropic and thigmotropic responses
Nastic response
nondirectional turgor movement in plants. turgor pressure is the pressure of the contents of the cell pressing against the cell wall and is regulated by the moving water into/out of the cell/vacuole. They are proportion to the stimuli
Most common nastic response
the opening and closing of stomata due to nastic movement of guard cells
Time frame (tropic vs Nastic)
nastic responses are immediate where as tropic responses occur over a long period of time
Reversibility (tropic Vs nastic)
Nastic responses are reversible, tropic responses are permanent
Response mechanism (nastic)
often rely on specialized joint-like thickening at the base of a leaf called a pulvinus
Pulvinus joint
consists of a vascular core (xylem and phloem) surrounded by a spongy flexible thin walled cell (two types, extensors and flexors)
Extensor vs flexor cells
extensors increase their turgor pressure during leaf ‘opening’ and decrease during ‘closing’. flexor cells do the opposite
How the response is affected (nastic)
response is affected due to active transport of sucrose and ions such as potassium and chloride. As these solutes are moved between extensors and flexors, water flows due to osmosis.
Photoperiodism
To ensure the best chance of survival and reproduction, plants need to be in sync with seasonal rhythms in temperature, precipitation, pollinator activity and sunlight
Stimuli of photoperiodism
Many stimuli could be used to detect the season, however, none are as reliable as the relative length of day and night (the photoperiodism)
Critical period
The critical duration of day/night required to stimulate or inhibit flowering
Long day plants
only flower is the day length exceeds their critical day length. This is generally around 14-16 hours. Long day plants generally flower in late spring and summer.
Short day plants
Only flower if the day length is less than their critical day length. generally, 8 - 10 hours. Theses plants generally flower in winter and autumn
Day neutral plants
photoperiod has no effect on day neutral plants. They instead generally inhibit flowering after reaching an age/development stage and /or in response to environmental cues such as a perios of sustained temperature.
taxes and kinesis
are simple innate movement in reponse to stimuli. This allows organisms to move to a favourable environment, giving them a survival advantage
taxes
directional movements, can be positive or negative
kinesis
non-directional to stimuli. instead the rate of movement changes in relation to the comfort zone of the organism. kineses are very common in single-celled organisms
Types of kinesis
orthokinesis and klinokinesis
Orthokinesis
changes the speed of movement proportional to the stimuli
Klinokinesis
Changes the rate of turning proportional to the level of stimuli
innate behaviour
those that occur instinctively due to genetics
learned behaviour
is developed overtime by modification of behaviour due to previous experience
Homing
an animals instinctive ability to navigate through unfamiliar territory and return to an original location using navigational cues.
When is homing need
when a species is displaced from their familiar territory during migration, mating or even simply during hunting or other regular activity
Navigational cues/methods
position of the sun, position of the star patterns (celestial navigation), orientation of Earth’s magnetic field, direction of ocean currents, scents and pheromones (olfaction) and sound
Navigating familiar terrain
many animals use this navigating, however, this is not considered a type fo homing as it only occurs in familiar territory
migration
The movement of animals from one region to another. True migration involves annual or season movements, or involves a major habitat change as a part of a species life
How is migration controlled/initatied
migration is genetically initiated by the environment. For migration to occur the advantages must outweigh the disadvantages
complete migration
all members of the population will migrate
Partial migration
some members of the population will migrate
differential migration
all members of the population migrate but not necessarily all at the same time of travelling the same distance. The differences are often due to age or sex
irruptive migration
members of the population do not migrate at all some years and do migrate other years
advantages of migration
new/more resources, constant food supply, find mates, greater genetic mixing, better breeding conditions, reduces predation, disease and parasitism, remain in an favourable climates, lead to colonisation of new areas
limitation of migration
uses lots of energy, animals can run out of energy before reaching their destination and die, risk of getting lost or eaten on the way
cues for migration
cues for migration involve complex interactions between internal and external stimuli
external stimuli (migration)
changes in weather, food and water supply and lenght of day
Internal stimuli (migration)
circannual rhythm (internal calendar), fat reserves, sex maturity
Rhythms
most environments on earth undergo predicatble fluctations due to the time of day, month (moon phase) and year. Animals have a survival adventage if they can predict, prepare and respond to these fluctuations
Keeping time
rhythms are maintained due to endogenous pacemakers (biological clocks) and exogenous zeitgebers (environmental cues)
Biological clocks
The bodies innate time keeping sytem. Occurs due to complex interaction of certain genes and proteins called a molecular clock. Pressent in almost all tissues in the body. The master clock is a group of 20,000 cells in the anterior hypothalamus called the SCN. This clock maintains circadian rhythm
entrainment
the process by which a biological clock is affected by environmental cues. Can occur at different levels of organisation from indivdual cells to whole organisms.
Jet lag
When the clock becomes out of sync with the environment you experience jet lag. Synchronization can only be restored with tim eand entrainment by zeitgebers. Humans have a natural biological clock that lasts 24hs and 11mins (± 16mins)
Suprachiasmatic nucleus (SCN)
the Suprachiasmatic Nucleus is located above the chiasma of your optic nerve adjacent to the pineal gland. When light enters the eye, special photoreceptor cells in the retina detect blue light and send a nerve impulse to the SCN. This signal increases the production and accumulation of CLOCK and BMAL1 proteins
CLOCK and BMAL1
The CLOCK and BMAL1 proteins also promote the transcription of period (PER) and cryptochrome (CRY) proteins. When in high abundance they form the PER/CRY complex. This inhibits both the secretion of melatonin and the transcription of more PER and CRY protein. This leads to wakefulness.
Diurnal
Active during the day. This provides many advantages for animals that have poor visibility in low light (like humans)
Nocturnal
Active during the night. Advantageous for avoiding predators and competition. In some areas it can be an adventage to avoid the heat.
Crepuscular
Active during twilight (dusk and dawn) adventageous for avoiding the heat while utilising light for activities such as hunting.
Actograms
diagrams/graphs to show animal activity patterns over a period, demonstarting the circadian rhythm of animals.
Purpose fo actograms
Shows a period when zeitgeber is present and a period when it is removed, there is useful for determining the relative influence of endogenous (internal) and exogenous (external) factors.
phase shift
When no zietgebers are present, the organism is said to be in a ‘free running’ period.If the activity pattern was entirely endogenous, it would remain relatively similar. However, in the absence of environmental cues the activity pattern is likely to drift. The amount of drift is the phase shift.
Levels of ecological organisation
1. organism, 2. population, 3. community, 4. ecosystem, 5. biome, 6. biosphere
population
all the same organisms
community
all the same species within an environment
ecosystem
all the biotic and abiotic factors within an environment
biome
a major biotic community characterised by dominant factors
Biosphere
the zone of all life on earth
interspecific relationships
the way different species within a community interact with each other eg, mutualistic, exploitative, commensalistic
mutualistic relationship
benefits both species
commensalistic relationship
benefits one species while not harming the other
exploitative
benefits one species while harming the other
competitive
harming both species
symbiosis
used to describe a close and long-term interaction between organisms of different species. Symbiotic relationships can be mutualistic commensalistic or parasitic
Radicle
positively gravitropic- is growing downwards
Plumule
negative gravitropism - grows away from gravity towards the light
PFR in long-day plants.
PFR promotes flowering in long-day plants. When PFR stays above the threshold due to short nights, flowering is able to occur. However, when nights are long too much PFR decays into PR so flowering is inhibited.
PFR in short-day plants.
Pfr inhibits flowering in short day plants. When Pfr drops below the threshold during long nights and converts to Pr, there is enough Pr to promote flowering. However, short day plants will not flower when the nights are too short, too much Pfr remains and inhibits flowering.
complex hierachies
has subordinate and affiliations within the overall group, such as mating pairs, and associations are constantly changing. there is an alpha male in overall charge
linear hierachies
when each animal is dominant over those below ir and submissive to those above it in the hierarchy. This is maintained by ritualised displays and aggresion.
advantages of hierarchies
lowers conflict and aggression as hierarchy is determined early in life during brief fights or threats and displays after which status is seldom contest except by newcomers.
adventages of living as a group
protection from the physical environment, great ability to detect predators, better defense, mates available, provision of learning opportunities for young.
disadvantages of living in a group
increased intraspecific competition, increased risk of contracting and spreading disease, access to mates may be restricted to only higher ranked individuals
what is the benefit of increased reproduction
increase the frequency of favourable alleles in the gene pool which increases chances of survival of individuals and increases genetic diversity resulting in an increased population size.
when using celestial navigation what must organisms also have?
a biological clock to take into account the rotation of the earth which causes the apparent movement of stars
how do auxins cause elongation of cells?
auxins loosen the cell walls which then take on more water and elongatie; therefore growing faster then the cells on the other side