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lime
this may be spread on pastures to raise soil pH
alfalfa
a common legume forage
lysine
the first limiting amino acid in horses
-only amino acid with known requirements
soybeans and soybean meal
animal sources (whey protein)
-grasses and many grains are generally low in lysine
forage
a horse should eat at least 1.5% of its body weight per day in ______, on a dry matter basis
volatile
microbial fermentation of fiber produces _____ fatty acids, which are used as an energy source by the horse
laminitis
overload of sugar and starch in the diet can cause this if it is not digested in the small intestine
esophagus
the long muscular tube between the horse’s mouth and its stomach
lignin
indigestible fiber that passes through the horse and is not utilized
fat
if we want to increase the body condition score of a horse, we could consider feeding a diet higher in this energy source
iodine
a micromineral found in higher concentrations in grasses than legumes
small intestine
site of protein digestion in the horse
stomach
because it is designed to be eating throughout the day, the horse’s ______ is relatively small, compared to some other species
net energy
the fraction of gross energy used for maintenance and production
soil sample
this should be performed every three years to determine soil pH and fertilizer needs
cecum
a large component of the hindgut in which forage is fermented by microbes
2:1
an appropriate calcium:phosphorus ratio would be _____
niacin
another name for vitamin B3
calcium
a macromineral important for bone strength and muscle contractions
glucose
a major form of carbohydrate stored in the muscle and liver as glycogen
vitamin A
a fat-soluble vitamin whose precursor, beta-carotene is found in green leafy plants
structural
fiber is an example of a ______ carbohydrate
energy
body condition scoring is a measure of the _____ status of a horse by determining how much fat cover it has
gall bladder
an organ in the digestive tract of a horse that humans have but horses do not
salt
a feedstuff that should always be available to horses
biotin
a B vitamin that is well-known for its role in hoof growth and repair
modified monogastrics
horses are _____ ______
-fermentation in the cecum
-able to utilize large amount of roughage in the diet
mouth
part of the body that performs prehension and ingestion
mastication
alteration of physical form
-reduces particle size, which improves digestibility
-greater number of chewing movements for roughage than concentrate
long
_____ stem forage promotes more even dental wear
99%
saliva is greater than _____ water, contains almost no digestive enzymes
10-12
how many liters of saliva does an average horse produce in a day?
bicarbonate
what does saliva contain that acts as a gastric pH buffer?
angle
the sharp ____ of the esophagus at the attachment to the stomach is the reason why horses cannot belch or vomit
7.5 to 15
how many liters does the average stomach hold?
non-glandular
the upper mucosal surface of the stomach is lined with ____ (squamous) epithelium
glandular
the lower region of the stomach is lined with ____ mucosa
hydrochloric acid, pepsin, bicarbonate, mucus
what 4 things does the stomach secrete 24/7
pepsin and lipase
what two enzymes begin to digest proteins and fatty acids in the stomach?
hind gut
where microbial fermentation of fiber and water absorption occurs
foregut
where enzymatic digestion of sugars, starch, most protein, fat, vitamins, minerals occurs
small intestine
site for digestion of many vitamins, some minerals
-relatively short (about 35% of the digestive tract)
-rapid passage (45 minutes to 4 hours)
-very little microbial fermentation of fiber
amino acids
pancreatic and SI enzymes digest protein into _____
fatty acids
pancreatic and SI enzymes digest fats into _____
glucose, simple sugars
pancreatic and SI enzymes digest soluble or hydrolysable CHO (sugars and starch) into ______
duodenum, jejunum, illeum
what are the three sections of the small intestine
villi
the presence of these in the small intestine help to increase the surface area
monosaccharides
simple sugars that are digested in the small intestine
oligosaccharides
short chain monosaccharides joined together, also sugars
-digested in the small intestine
polysaccharides
usually starch and cellulose
-long chains of 100s to 1000s of monosaccharides
starch
subject to enzymatic digestion in the small intestine
cellulose
requires microbial cellulase in the hindgut
small intestine
sugars and starches are non-structural carbohydrates and are digested in the _____
hindgut
dietary fibers are structural CHO and are digested in the ____
hindgut
term for the combination of the cecum and colon (large and small)
-largest portion of the GI tract (~60%)
-rate of passage is 50-60 hours
-no mucosal enzymes
-where water is absorbed
cecum
one of the most important parts of the GI tract!
-8 gallon volume
-microbial digestion by bacteria, fungi and protozoa
create volatile fatty acids that horse can use as an energy source
B and K
what vitamins does a horse synthesize themselves in the cecum?
large colon
10-12 feet in length, 21.5 gallon capacity
-nutrient absorption, continued fermentation
small colon
10-12 feet in length, 5 gallon capacity
-water and mineral absorption, fecal ball formation
water, energy, protein, vitamins, minerals
what are the 5 main nutrient classes
water
the most important nutrient, often overlooked
-responsible for
maintaining body fluid balance
digestive functioning
gastrointestinal health
62-68%
total body water in adult horses is about ____ of body weight
drinking, moisture in feed
2 examples of direct water intake
metabolism of fats, CHO, protein
what is an example of indirect water intake
urination, defecation, respiration, lactation, cutaneous
5 examples of water loss
water-holding
fiber has a high _____ capacity
urinary
horses consuming excess protein, potassium, or sodium will have an increased renal solute load, and thus higher ____ water loss
digestible energy
gross energy minus feces energy
metabolized energy
digestible energy minus urine and methane energy
net energy
metabolized energy minus heat increment
-used for maintenance and production
calorie
amount of heat produced by oxidation that raises one gram of water one degree celcius
1000 kcal
1 megacalorie equals ____ kcal
-used for horses
-not included on feed labels
digestible energy
total digestible nutrients
-affected by gross energy content of feeds
-can be determined from feeding trials
-also can be determined from prediction equations from the NRC
BCS 1
Poor. Animal extremely emaciated. No fatty tissue can be felt. Bone structure very noticeable. Ribs projecting prominently. Shoulder is sharp and protruding. Tailhead, point of hip and point of buttock projecting prominently.
BCS 2
Very Thin. Animal emaciated. Ribs prominent. Noticeable bone structure. Vertebrae prominent. Neck is obviously thin. Shoulder is noticeable. Ribs easily discernible. Points of hip and buttock prominent. Tailhead very prominent.
BCS 3
Thin. Slight fat cover over ribs. Ribs easily discernible. Neck, withers and shoulder accentuated. Hip is rounded with some filling, but easily discernible. Point of buttock slightly distinguishable. Tailhead prominent, but individual vertebrae cannot be visually identified.
BCS 4
Moderately Thin. Faint outline of ribs with some fat filling between ribs. Negative crease along back (backbone sticks up). Neck, shoulder and withers slightly thin. Point of hip still discernable. Point of buttock rounded and not distinguishable. Tailhead slightly prominent.
BCS 5
Moderate. Ribs not visible but are easily felt. Backbone is level. Shoulder blends smoothly into body. Withers rounded. Hip and buttock rounded. Fat around tailhead beginning to feel spongy, and tailhead blends smoothly with rump.
BCS 6
Moderately Fleshy. Fat beginning to be deposited. Fat over ribs feels spongy. Ribs are not visible, but can be felt with slight pressure. Slight positive crease (gutter appearance) may appear down backbone. Shoulder blends smoothly into body. Withers rounded. Fat beginning to be deposited around tailhead.
BCS 7
Fleshy. Fat deposited along neck, withers, area behind shoulder and on either side of tailhead. Neck may begin to become cresty. Individual ribs can be felt with pressure, but noticeable filling between ribs. Fat is beginning to fill in the flank. Back has positive crease (gutter appearance).
BCS 8
– Fat. Fat deposited at withers, area behind shoulders, and along either side of tailhead. Thickened and somewhat cresty neck. Difficult to feel ribs. Positive crease (gutter appearance) down back. Flank is nearly filled flush. Fat deposited along inner buttocks. Fat around tailhead feels spongy
BCS 9
Very Fat. Bulging fat. Patchy fat appearing over ribs. Difficult to palpate ribs. Area behind shoulder is filled with fat. Obvious positive crease (gutter appearance) down back. Fat along inner buttocks may rub together. Flank filled in flush. Tailhead appears sunken due to fat deposits along either side
body condition index
method of assessing body fat similar to BMI in humans
-not as subjective as BCS
one month
usually takes at least _____ to increase or decrease a BCS score
carbohydrates, fat, protein
where do horses get DE
-the energy containing nutrients
-energy from these nutrients can be
immediately available after a meal
stored for future use
ATP
major source of chemical energy used in cells
-adenosine triphosphate
-generated from metabolism of carbs, fats, and proteins
glucose
the major form of CHO used to generate ATP
-can come from circulation, or from body stores
-stored primarily in skeletal muscle and liver
fat
the most abundant energy source in the body
-broken down into fatty acids, which are then broken down to generate ATP
long chain fatty acids
come from food digestion or from stored fat (adipose tissue) in the form of triglycerides
short chain fatty acids
come from bacterial fermentation of structural carbohydrates in the hindgut
-aka volatile
-can meet most of energy requirements at maintenance
-can also be metabolized into long-chain fatty acids or glucose
carbohydrates
the most common form of energy in equine diets
-come from forages and concentrates
structural
type of CHO that consists of fiber
non-structural
type of CHO that consists of sugars, starches and pectins (sometimes a soluble fiber)
amylase
the enzyme that is missing in the small intestine to be able to break down fiber
glycogen
the storage form of glucose in liver and muscle
fiber
the most important ingredient in equine diets
-usually provides sufficient energy required for maintenance
crude fiber
crude assay related to structural CHO in the feed
-on feed labels
-somewhat indistinct
-most of cellulose, but only part of lignin
-always underestimates actual fiber content of a forage
-generally the higher the CF, the lower the DE and digestibility it will have
pectin
a water-soluble fiber that is found in the cell walls of fruits, tubers, and stems of plants
-in presence of low pH, these turn into a gell
neutral detergent fiber
percentage of the plant material in the forage made up of structural fibers
-higher it is the lower the intake, more wastage
acid detergent fiber
percentage of the plant material in the forage that is difficult for the horse to break down
-cellulose and lignin
-higher it is, the lower the digestibility and lower nutrient availability
water soluble
this kind of CHO includes simple sugars, disaccharides, oligosaccharides, and some polysaccharides