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population
A group of individuals of a single species living in the same general area.
density
The number of individuals per unit area or volume.
dispersion
The pattern of spacing among individuals within the boundaries of the population.
mark-recapture method
A sampling technique used to estimate the size of wildlife populations.
immigration
The influx of new individuals from other areas.
emigration
The movement of individuals out of a population and into other locations.
clumped
A pattern of dispersion in which individuals are aggregated in patches.
uniform
An evenly spaced pattern of dispersion.
territoriality
The defense of a bounded physical space against encroachment by other individuals.
random dispersion
Unpredictable spacing; the position of each individual in a population is independent of other individuals.
Demography
The study of the key characteristics of populations and how they change over time.
life table
A summary of the survival and reproductive rates of individuals in specific age-groups within a population.
cohort
A group of individuals of the same age.
survivorship curve
A plot of the proportion or numbers in a cohort still alive at each age.
exponential population growth
The growth of a population in an ideal, unlimited environment.
intrinsic rate of increase
The per capita rate at which an exponentially growing population increases in size at each instant in time.
carrying capacity
The maximum population size that a particular environment can sustain.
logistic population growth
A model where the per capita rate of population growth approaches zero as the population size nears the carrying capacity (K).
life history
The traits that affect an organism's schedule of reproduction and survival.
semelparity
A "one-shot" pattern of big-bang reproduction.
iteroparity
Repeated reproduction over an organism's lifetime.
K-selection
Selection for traits that are advantageous at high population densities.
r-selection
Selection for traits that maximize reproductive success in uncrowded environments (low densities).
density independent
A birth rate or death rate that does not change with population density.
density dependent
A death rate that increases with population density or a birth rate that falls with rising density.
population dynamics
Population fluctuations from year to year or place to place.
metapopulation
A number of local populations linked by immigration and emigration.
demographic transition
The movement from high birth and death rates toward low birth and death rates in human populations.
age structure
The relative number of individuals of each age in the population.
infant mortality
The number of infant deaths per 1,000 live births.
life expectancy at birth
The predicted average length of life at birth.
ecological footprint
The aggregate land and water area required by each person, city, or nation to produce all the resources it consumes and to absorb all the waste it generates.
community
A group of populations of different species living in close enough proximity to interact.
interspecific interactions
Ecological interactions between individuals of different species in a community.
competition
A -/- interaction that occurs when individuals of different species each use a resource that limits the survival and reproduction of both.
competitive exclusion
The local elimination of the inferior competitor due to two species competing for the same limiting resources.
ecological niche
The specific set of biotic and abiotic resources that an organism uses in its environment.
resource partitioning
The differentiation of niches that enables similar species to coexist in a community.
character displacement
The tendency for characteristics to diverge more in sympatric (overlapping) than in allopatric (geographically separate) populations of two species.
exploitation
Any type of +/- interaction in which individuals of one species benefit by feeding on (and thereby harming) individuals of the other species.
Predation
A +/- interaction in which an individual of one species (predator) kills and eats an individual of the other species (prey).
aposematic coloration
Warning coloration that warns predators that an animal has effective chemical defenses.
Cryptic coloration
Camouflage that makes prey difficult to see.
Batesian mimicry
When a palatable or harmless species mimics an unpalatable or harmful species to which it is not closely related.
Müllerian mimicry
When two or more unpalatable species resemble each other.
Herbivory
An exploitative (+/-) interaction in which an organism eats parts of a plant or alga.
Parasitism
A +/- exploitative interaction in which one organism derives its nourishment from another organism, which is harmed.
parasite
The organism that derives nourishment from its host in a +/- interaction.
host
The organism that is harmed in a parasitic interaction.
endoparasites
Parasites that live within the body of their host.
ectoparasites
Parasites that feed on the external surface of a host.
positive interactions
A +/+ or +/0 interaction between members of two species in which at least one individual benefits and neither is harmed.
Mutualism
A +/+ interaction that benefits individuals of both of the interacting species.
commensalism
An interaction that benefits the individuals of one of the interacting species but neither harms nor helps the individuals of the other species (+/0).
species diversity
The variety of different kinds of organisms that make up the community.
species richness
The number of different species in a community.
relative abundance
The proportion each species represents of all individuals in the community.
Shannon diversity index (H)
An index of community diversity based on species richness and relative abundance.
biomass
The total mass of all organisms in a habitat.
introduced species
Organisms that humans have moved to regions outside of the species' native range.
trophic structure
The feeding relationships between organisms in a community.
food chain
The transfer of chemical energy from its source in plants and other autotrophs through herbivores to carnivores and eventually to decomposers.
trophic level
The position an organism occupies in a food chain.
food web
A group of food chains linked together to show complex trophic interactions.
energetic hypothesis
Suggests that the length of a food chain is limited by the inefficiency of energy transfer along the chain.
Foundation species
Species that have strong effects on their communities as a result of their large size or high abundance.
keystone species
Species that are not usually abundant but exert strong control on community structure by their pivotal ecological roles.
ecosystem engineers
Species that create or dramatically alter their environment.
bottom-up control
When the abundance of organisms at each trophic level is limited by nutrient supply or the availability of food at lower trophic levels.
top-down control
When the abundance of organisms at each trophic level is controlled by the abundance of consumers at higher trophic levels.
disturbance
An event, such as a storm, fire, or human activity, that changes a community by removing organisms from it or altering resource availability.
nonequilibrium model
Describes most communities as constantly changing after a disturbance, rather than being at a stable balance.
intermediate disturbance hypothesis
States that moderate levels of disturbance foster greater species diversity than do high or low levels of disturbance.
ecological succession
The gradual process by which the species composition of a community changes and develops over time after a disturbance.
primary succession
Ecological succession that begins in a virtually lifeless area where soil has not yet formed.
secondary succession
The recolonization of an area after a major disturbance has removed most but not all of the organisms in a community.
evapotranspiration
The evaporation of water from soil and plants; used as a measure of climate impacts on species richness.
species-area curve
The concept that, all other factors being equal, the larger the geographic area of a community, the more species it has.
pathogens
Disease-causing microorganisms, viruses, viroids, or prions.
zoonotic pathogens
Pathogens that are transferred to humans from other animals.
vector
An intermediate species, such as a tick or mosquito, that transfers a zoonotic pathogen to a host.
ecosystem
The sum of all the organisms living in a given area and the abiotic factors with which they interact.
law of conservation of mass
A physical law stating that matter, like energy, cannot be created or destroyed.
primary producers
The trophic level that ultimately supports all others; consists of autotrophs.
primary consumers
Herbivores, which eat plants and other primary producers.
secondary consumers
Carnivores that eat herbivores.
tertiary consumers
Carnivores that eat other carnivores.
decomposers
Heterotrophs (consumers) that get their energy from breaking down detritus.
detritus
Nonliving organic material, such as the remains of dead organisms, feces, and fallen leaves.
primary production
The amount of light energy converted to chemical energy by autotrophs during a given time period.
gross primary production (GPP)
Total primary production in an ecosystem.
Net primary production (NPP)
Gross primary production minus the energy used by the primary producers for their own cellular respiration.
net ecosystem production (NEP)
A measure of the total biomass accumulation by both producers and consumers during a given period.
limiting nutrient
The element that must be added for primary production to increase in an ecosystem.
eutrophication
A process by which nutrients, particularly phosphorus and nitrogen, become highly concentrated in a body of water, leading to increased growth of organisms such as algae.
secondary production
The amount of chemical energy in consumers' food that is converted to their own new biomass during a given period.
production efficiency
The percentage of energy stored in assimilated food that is used for growth and reproduction, not respiration.
trophic efficiency
The percentage of production transferred from one trophic level to the next (averaging about 10%).
biogeochemical cycles
Nutrient cycles that involve both biotic and abiotic components of ecosystems.
bioremediation
The use of organisms—usually prokaryotes, fungi, or plants—to detoxify polluted ecosystems.