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Darwin's Revolution (1859)
Introduced the idea that species change over time through natural selection.
Descent with Modification
Species evolve from common ancestors, accumulating adaptations over generations.
Aristotle's view on species
Species are fixed and arranged by complexity.
Carl Linnaeus
Developed binomial nomenclature and classification systems.
Binomial Nomenclature
Classification system in which each species is assigned a two-part scientific name
Classification systems
Specific sets of objective criteria, such as offense history, previous experience in the justice system, and substance abuse patterns, applied to all inmates to determine an appropriate classification.
Georges Cuvier
Proposed catastrophism based on fossil changes over time.
James Hutton & Charles Lyell
Proposed uniformitarianism, implying the Earth is shaped by gradual processes.
Jean-Baptiste Lamarck
Proposed use/disuse and inheritance of acquired traits.
HMS Beagle Voyage
Darwin's voyage where he noted similarities between island and mainland species.
Galápagos Finches
Showed variation in beak shape linked to diet and environment.
Natural Selection
Process where individuals with advantageous traits survive and reproduce more.
Natural Selection: Key Observations
Traits vary within populations, and more offspring are produced than survive.
Natural Selection: Key Inferences
Favorable traits increase survival and accumulate over generations.
Soapberry bugs (evolution example)
Beak length adapted rapidly to changes in fruit size.
MRSA (evolution example)
Demonstrates rapid evolution of antibiotic resistance in bacteria.
Anatomical Homology
Similar structures from common ancestry, like mammalian forelimbs.
Molecular Homology
Shared genes and genetic code across different species.
Vestigial structures
Remnants of ancestral traits that serve little or no function.
Fossil Record
Shows evolutionary transitions, such as land-to-sea in cetaceans.
Biogeography
Study of species distribution influenced by continental drift.
Adaptation
An inherited trait that increases an organism's fitness.
Artificial Selection
Human-driven breeding for desired traits.
Convergent Evolution
Independent evolution of similar traits in unrelated groups.
Evolutionary Tree
Diagram showing evolutionary relationships and common ancestry.
Unit of Evolution
Populations evolve over time; individuals do not.
Origin of traits in selection
Natural selection acts on existing variation; it cannot create new traits.
Role of environment in selection
The environment determines which traits are favorable.
Microevolution
Change in allele frequencies in a population over generations.
Three mechanisms of microevolution
Natural selection, genetic drift, and gene flow.
Natural Selection
Differential survival and reproduction
Genetic Drift
Random changes in allele frequencies
Gene Flow
Movement of alleles between populations
Sources of genetic variation
Mutations, gene duplication, and sexual reproduction.
Three sources of allele recombination in sexual reproduction
Crossing over, independent assortment, and fertilization.
Phenotypic vs. Genotypic variation
Phenotypic variation affects observable traits; genotypic variation measures molecular differences.
Five conditions for Hardy-Weinberg equilibrium
No mutations, random mating, no natural selection, extremely large population, no gene flow.
Hardy-Weinberg equilibrium outcome
If all conditions are met, the population is not evolving.
Genetic Drift
Random changes in allele frequencies, having stronger effects in small populations.
Founder Effect vs. Bottleneck Effect
Founder: few individuals start a population. Bottleneck: sudden reduction in population size.
Gene Flow
Movement of alleles between populations, tending to reduce genetic differences between them.
Target of Natural Selection
Natural selection acts on phenotypes, not genotypes.
Relative fitness
The contribution an individual makes to the next generation's gene pool.
Three modes of natural selection
Directional (one extreme), disruptive (both extremes), and stabilizing (intermediate traits).
Intrasexual vs. Intersexual selection
Intrasexual: competition within one sex. Intersexual: mate choice.
Good Genes Hypothesis
The hypothesis that females prefer traits signaling genetic quality.
Balancing Selection
Selection that maintains multiple alleles in a population.
Types of balancing selection
Frequency-dependent selection (fitness depends on how common a trait is) and heterozygote advantage (heterozygotes have higher fitness).
Speciation
The process by which one species splits into two or more species.
Speciation Bridge
Bridges microevolution (allele frequency changes) and macroevolution (broad evolutionary patterns).
Biological Species Concept
Species are groups of populations that interbreed, produce viable, fertile offspring, and are reproductively isolated.
Reproductive Isolation
Biological barriers that prevent gene flow between species.
Habitat Isolation
Prezygotic barrier where species occupy different environments and rarely encounter each other.
Temporal Isolation
Prezygotic barrier where species breed at different times of day, seasons, or years.
Behavioral Isolation
Prezygotic barrier where unique courtship rituals or mating behaviors prevent interbreeding.
Mechanical Isolation
Prezygotic barrier where morphological differences prevent successful mating.
Gametic Isolation
Prezygotic barrier where sperm of one species cannot fertilize eggs of another.
Reduced Hybrid Viability
Postzygotic barrier where genes of parents impair the hybrid's development or survival.
Reduced Hybrid Fertility
Postzygotic barrier where hybrids develop but are sterile (cannot produce offspring).
Hybrid Breakdown
Postzygotic barrier where first-generation hybrids are fertile, but their offspring are feeble or sterile.
Allopatric Speciation
Speciation that occurs when geographic separation restricts gene flow between populations.
Sympatric Speciation
Speciation that occurs in populations that live in the same geographic area.
Drivers of Sympatric Speciation
Polyploidy, sexual selection, and habitat differentiation.
Hybrid Zones
Regions where members of different species meet and mate, producing mixed offspring.
Hybrid Zone: Reinforcement
Strengthening of reproductive barriers, leading to a decrease in hybrid formation.
Hybrid Zone: Fusion
Weakening of reproductive barriers, causing the two species to merge into one.
Hybrid Zone: Stability
Continued production of hybrid individuals without fusion or reinforcement.
Punctuated Equilibria
Periods of apparent evolutionary stasis punctuated by sudden change.
Gradualism
Slow, steady evolutionary change over long periods of time.
Genetics of Speciation
Can involve a single gene or multiple genes to achieve reproductive isolation.
Early Earth Conditions
Hostile environment with volcanic activity, toxic atmosphere, and no oxygen, where simple organic molecules formed.
Hydrothermal Vents Hypothesis
Theory that life began near ocean-floor vents providing heat, minerals, and chemical-rich environments.
Role of RNA in early life
Simple molecules formed RNA, which could store information, catalyze reactions, and self-replicate.
Protocells
Primitive cell-like structures formed by molecules enclosed in membranes that could grow and divide.
Evolution of early molecular systems
Natural selection favored more efficient and stable molecular systems, leading to diverse life.
Macroevolution
Evolutionary changes above the species level.
Strata
Sedimentary rock layers where most fossils are found.
Radiometric dating
Method using isotope decay to determine fossil age.
Bias in the fossil record
Favors widespread, long-lived species with hard parts.
Four eons of Earth's history
Hadean, Archaean, Proterozoic, and Phanerozoic.
Eras of the Phanerozoic eon
Paleozoic, Mesozoic, and Cenozoic.
Stromatolites
Fossilized structures representing the first life 3.5 billion years ago.
Oxygen revolution
Atmospheric oxygen accumulation occurring 2.7 to 2.4 billion years ago.
First eukaryotes
Appeared in the fossil record 1.8 billion years ago.
First multicellular eukaryotes
Appeared in the fossil record approximately 1.2 billion years ago.
Cambrian explosion
Rapid diversification of animal phyla about 535 million years ago.
Endosymbiosis theory
Mitochondria and plastids originated from engulfed prokaryotes.
Evidence for endosymbiosis
Similarities in DNA, ribosomes, and reproduction between organelles and bacteria.
Synapsids
Group of organisms from which mammals evolved.
Mammalian skeletal evolution
Gradual changes in jaw and ear bones.
First land colonizers
Prokaryotes, which arrived on land around 3.2 billion years ago.
Key animal land colonizers
Arthropods and tetrapods.
Pangaea
Supercontinent that formed 250 million years ago.
Effects of continental drift
Alters climate, habitats, and drives speciation.
Permian extinction
Mass extinction 252 million years ago caused by volcanic activity.
Cretaceous extinction
Mass extinction 66 million years ago caused by meteorite impact.
Adaptive radiation
Rapid evolution of species to fill vacant ecological niches.
Heterochrony
Evolutionary change in the timing or rate of development.
Homeotic genes (Hox genes)
Genes that control the spatial layout of body plans.
Long-term evolutionary trends
Shaped by environmental change, extinction, and evolutionary innovation.