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Macroevolution
Evolutionary change above the species level
Role of Direct Observations for collecting evidence
Evolutionary changes are documented by overwhelming amounts of biological studies
especially effective for studies on organisms with short life spans
Homologous Traits
Homology
Comparative embryology
Comparative anatomy
Molecular homology
Homology
Character traits that evolved from a common ancestral structure
Comparative embryology
homologous features visible during embryonic development
More general features appear early in development, acting as supporting evidence for shared ancestry
Comparative anatomy
homologous features visible on individuals after birth
Homology of limbs in vertebrates
Vestigial structures as evolutionary remnant of ancestors (evolution removes original function)
Molecular homology
homologies are demonstrated by similarities in genetic make-up inherited from a common ancestor
Degrees of similarity is determined by how long ago they diverged from their common ancestor
Fossils
remains of species in Earth’s history
Fossils are indisputable evidence that ancestral species existed in Earth’s history.
Can document “in-between” stages of evolution
Issues with Fossil Record
Fossil record is incomplete and biased:
Some biological materials are easier to fossilise
Some species live in environments that favor preservations.
Fossil Record Contributions to Study
Biology of extinct species
Origin and diversification of some clades (e.g. the clade of birds branched off the clade of dinosaurs)
Relative branching order among clades (e.g. the clade of amphibians first branched off, then the clade of amniotes, etc.)
Tracking of evolution over a long, long time
Calibration of geological time for molecular study
Biogeography
The study of past and present geographic distribution of species
Biogeography Contributions to Study
Assist in reconstruction of historical paths that animals traversed (e.g. how Canada geese migrate)
Document distribution of phenotypic and genotypic variations.
Monitor interactions between extant species that may lead to evolutionary changes (e.g. interactions between wolves and moose on Isle Royale National Park)
Endemic Species
Biological taxa that are native to a specific geographic area, and not found in other regions (often found in remote location)
Selection Pressure in Endemic Species
Populations of endemic species would experience selection pressure different from that experienced by populations with wider distribution
Over time, endemic species may accumulate sets of genotypic and phenotypic variations that lead to speciation events