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How Should We Feed Animals?
The Main Question
What to feed?
How much of each nutrient to provide?
Traditional Approach
Tradition / know-how → based on experience and established practices
What we always fed → continue using familiar feeds
Animal-Driven Approach
Let the animal choose → allow the animal to select what it eats
Availability-Based Approach
Feed what we have → use whatever feed is available
Scientific Approach
Systematically + scientifically → determine what + how much to feed based on scientific evidence
Nutrition / nutritional sciences → science that drives feeding decisions

Nutrition: Definitions
American Medical Association (AMA)
Human-focused definition
Nutrition → science of food, nutrients + substances, including their actions, interactions + balance
Relates to health + disease
Includes processes by which the organism:
Digests food substances
Absorbs food substances
Transports food substances
Utilizes food substances
Excretes food substances
Kellems and Church (2002)
Animal production-focused definition
Nutrition → series of processes by which an animal takes in + assimilates feed components
Supports:
Growth
Milk production
Fiber production
Replacement of worn or injured tissues
Key Difference
AMA → focuses on nutrition + health/disease
Kellems & Church → focuses on nutrition + animal production

Human Nutrition vs. Animal Nutrition?
Important Context
My old boss’ view (1990s) → presented for discussion
The lecturer does not agree with this view
Human Nutrition
Called “Social Dietetics?”
Focuses on social + medical aspects
Goal → minimize risks to health
Animal Nutrition
Called “True science, serious + systematic?”
Focuses on maximizing growth + efficiency of livestock
Goal → minimize cost
Main Comparison
Human nutrition → health + social/medical concerns
Animal nutrition → growth + efficiency + cost

The Boundary Between Animal and Human Nutrition
Changing Boundary
The boundary between animal nutrition and human nutrition is no longer clearly defined
The two fields increasingly overlap
Why This Matters
Animal nutrition is not only about growth, efficiency + cost
Human nutrition is not only about health + social/medical aspects
Both can involve health, nutrition, food quality + well-being

Similarities Between Human and Animal Nutrition Research
Human Nutrition Group’s Mission
Advance knowledge about the role of nutrition in human health
What They Study
Whole foods
Individual nutrients
Non-nutritive food components
Their roles at the:
Whole body level
Cellular level
Molecular level
Research Groups
Studied in different population groups
Also studied using model systems
Food Choices
Environmental determinants → factors affecting food choices
Social determinants → social factors affecting food choices
Individual determinants → individual factors affecting food choices
Examines food choices + eating patterns
Animal Nutrition Research
This description could also be applied to much of the animal nutrition research carried out around the world

Nutrition and Feeding of Animals
Long History
Nutrition + feeding of animals → systematically approached for millennia
Ancient Egyptians
Developed a system to determine how much feed resources different types of cattle would need
18th + 19th Century
Animal nutrition became a focus of scientific efforts
Important work in:
Italy
France
United Kingdom
Germany
20th Century
Animal nutrition research became especially important in the USA

The Ahmes Papyrus – 1650 B.C.E.
Historical Record
Ahmes Papyrus → dates to 1650 B.C.E.
Shows the long history of systematic knowledge in ancient Egypt
Time Period
1650 B.C.E. → Ancient Egypt

Lavoisier and Laplace: Caloric Measurements on Animals and Humans
Time Period
1776–1791
17th + 18th Century
Part of the “Chemical Revolution”
Studies
Conducted caloric measurements on animals + humans
Performed complete mass + energy balance
Historical Significance
Important early work in understanding energy balance in animals and humans
Their work is associated with one of the best museums in Paris

Protein as a Nutrient
17th + 18th Century
Part of the “Chemical Revolution”
Protein
Protein → called “animal substance”
Considered the sole nutrient

Early Nutrient Requirements
Wolff, 1874
Requirements given in kg per day per 1,000 kg live-weight
Oxen at rest
Protein → 0.7 kg
Carbohydrate → 8.0 kg
Fat → 0.15 kg
Milch cow
Protein → 2.5 kg
Carbohydrate → 12.5 kg
Fat → 0.40 kg
Haecker, 1903
Maintenance requirements given in kg per day per 1,000 kg live-weight
Protein → 0.7 kg
Carbohydrate → 7.0 kg
Fat → 0.10 kg
Milk Production
Requirements also given in kg per day per kg milk
2.5% milk fat → Protein 0.0362 kg, Carbohydrate 0.164 kg, Fat 0.0124 kg
2.6% milk fat → Protein 0.0369 kg, Carbohydrate 0.167 kg, Fat 0.0126 kg
2.7% milk fat → Protein 0.0376 kg, Carbohydrate 0.171 kg, Fat 0.0128 kg
7.0% milk fat → Protein 0.0677 kg, Carbohydrate 0.313 kg, Fat 0.0229 kg
Memory Trick
Wolff = 1874 → oxen + milch cow
Haecker = 1903 → maintenance + milk production
More milk fat → higher nutrient requirements

Requirements of Animals – Information Sources
NRC – National Research Council
NRC → National Research Council
Provides information on the nutrient requirements of animals
Used as a reference for determining what nutrients animals need

Requirements of Animals – Authoritative Sources
Other Authoritative Documents
Published by different organizations
Provide information on requirements of animals
National or Regional Resources
Resources developed at a national or regional level

Poultry Nutritional Specifications – Public Guidelines
Major Public Guidelines
National Research Council (NRC)
INRAE-CIRAD-AFZ
Centraal Veevoederbureau (CVB)
Brazilian Tables (CBPA)
Agricultural Research Council (ARC)
Swedish Poultry Council (SPC) / AFZ
New Zealand Poultry Feed Tables (NZWG)
Foundation for Animal Nutrition (FEDNA)
Australian Poultry Standards
Chinese Poultry Nutrient Requirements → China Feed Database / NRC-Chinese adaptation
Japanese Poultry Feeding Standards (JPFS)
Korean Feeding Standards for Poultry (KFSP)
Guidelines from Major Broiler Breeders
Cobb Breeders
Ross Breeders
Hubbard Breeders
Arbor Acres Breeders
Hy-Line / ISA Layers

Genetic Potential & Feeding Management
Different Breeds
Different breeds have different genetic milk yield potential
This means their genetics determine how much milk they are capable of producing
Individual Animals
All animals have different genetic potential for:
Milk
Fat
Protein
Etc
Feeding Challenge
Different animals have different production potentials
Different production potentials mean different nutritional needs
Key Question
How do you manage the feeding of different animals with different production potentials and therefore different nutritional needs?

Animal Performance & Feeding
Constant Genetic Improvement
There are constant improvements in the performance of farm animals
Animals are rapidly “changing” over time
Feeding Must Change
As animals change over time, their feeding must also change
Feeding needs to keep up with improvements in animal performance
Different Genetic Potentials
Animals have different genetic potentials
Animals with different genetic potentials must be fed differently

Factorial Model – NRC, 1978
Basic Idea
Factorial Model → used to determine the animal’s nutrient requirements
Developed by NRC, 1978
Looks at the nutrients supplied by feed and the nutrients required by the animal
The goal is to achieve a BALANCE
Crude Protein Balance
FEED
Crude protein
↓
ANIMAL
Urine N
Scurf N → nitrogen lost from the skin
Metabolic fecal N → nitrogen lost in feces from the animal’s own metabolism
Protein yield
Net Energy Balance
FEED
Net energy
↓
ANIMAL
Dry matter intake
Body weight
Milk yield
Fat %
Key Idea
Feed nutrients → Animal needs → BALANCE
Crude protein → balanced against nitrogen losses + protein yield
Net energy → related to dry matter intake, body weight, milk yield + fat %

Absorbed Protein Model – NRC, 1988
Basic Idea
Absorbed Protein Model → determines whether the animal receives enough absorbed protein and net energy
Developed by NRC, 1988
Uses BALANCE+ and BALANCE− to compare what is supplied by the FEED with what the ANIMAL requires
FEED
Protein
Undegradable protein → protein that is not degraded in the rumen
Degradable protein → protein that is degraded in the rumen
Energy
Net energy
↓
ANIMAL
Protein Balance
UIP → undegradable intake protein
DIP → degradable intake protein
Absorbed protein
Endogenous N → nitrogen coming from the animal itself
Scurf N → nitrogen lost from the skin
Metabolic fecal N → nitrogen lost in feces from the animal’s own metabolism
Bacterial protein
Protein yield
Net Energy Balance
Net energy
Dry matter intake
Body weight
Milk yield
Fat %
Key Idea → Feed protein + energy → absorbed protein + net energy → animal production

Mechanistic Digestion Model – CNCPS, 1992 + NRC, 2001
Basic Idea
Mechanistic Digestion Model → describes how different feed components are digested and used by the animal
CNCPS, 1992
NRC, 2001
The model follows nutrients through digestion to determine metabolizable protein and metabolizable energy
The nutrients are then compared with what the ANIMAL needs to maintain BALANCE
FEED
Protein Components
NPN → non-protein nitrogen
B1 protein
B2 protein
B3 protein
Other Feed Components
Sugar
Starch
Fat
Avail fibre → available fibre
↓
DIGESTION
Ammon → ammonia
NSC bacteria → bacteria using non-structural carbohydrates
SC bacteria → bacteria using structural carbohydrates
↓
ANIMAL
Protein Balance
Metabolizable protein
Metabolic fecal N
Endogenous scurf N
Protein yield
Energy Balance
Metabolizable energy
Body weight
Milk yield
Protein %
Fat %
Key Idea → Feed components → Digestion → Metabolizable protein + metabolizable energy → Animal balance

Nutrient Requirement Models – Basics & Differences *FROM CHAT
1⃣ Factorial Model – NRC, 1978
Basic Idea
Looks at the nutrients going into the animal from the feed
Compares them with the nutrients used or lost by the animal
Goal → achieve a BALANCE
Main Nutrients
Crude protein
Net energy
What it considers
Urine N
Scurf N
Metabolic fecal N
Protein yield
Dry matter intake
Body weight
Milk yield
Fat %
Main Concept → Feed nutrients → animal needs/losses → balance
2⃣ Absorbed Protein Model – NRC, 1988
Basic Idea
Focuses more specifically on protein absorbed by the animal
Separates feed protein into different types based on how they behave during digestion
Also considers net energy
Main Nutrients
Undegradable protein
Degradable protein
Net energy
What it considers
UIP
DIP
Absorbed protein
Endogenous N
Scurf N
Metabolic fecal N
Bacterial protein
Protein yield
Dry matter intake
Body weight
Milk yield
Fat %
Main Concept → Feed protein → digestion + absorption → absorbed protein → animal production
3⃣ Mechanistic Digestion Model – CNCPS, 1992 + NRC, 2001
Basic Idea
Gives a more detailed description of what happens during digestion
Breaks feed into multiple components and follows how they are digested and used
Determines metabolizable protein and metabolizable energy
Main Feed Components
NPN
B1, B2, B3 protein
Sugar
Starch
Fat
Available fibre
What it considers
Ammonia
NSC bacteria
SC bacteria
Metabolizable protein
Metabolic fecal N
Endogenous scurf N
Protein yield
Metabolizable energy
Body weight
Protein %
Milk yield
Fat %
Applying Systematic Approaches to Animal Nutrition
Back to Basics: Intake ↑, MP:ME Dialled In, Results Followed
Systematic approaches → based on scientific principles and current knowledge
Presented by Maria Antonieta Puerto, MSc Animal Science, McGill University
Main approach → focus on intake, MP:ME, and then follow the results
Results
Herd improved from:
36 kg milk/cow/day → 43 kg milk/cow/day
1.5 kg fat/cow/day → 1.9 kg fat/cow/day
What Changed?
Make DMI the North Star
DMI → dry matter intake
Stopped guessing intakes and started measuring them
Feed access was improved with:
24/7 push-ups
Consistent TMR
Correct chop length
Correct particle size
Cows moved from 22 kg to 25 kg DM/day
The new ration's fuel → the increased DMI
Forage Quality
Forage quality was considered non-negotiable
Selecting forages with higher fiber digestibility → more nutrients available per bite
Starch Digestibility
Starch digestibility is not static → it can change depending on several factors
Factors affecting total-tract starch digestibility:
Hybrid
Maturity
Moisture
Kernel processing score
Particle size
Storage time
Meta-analyses show that improving kernel processing and particle size increases total-tract starch digestibility
Increased starch digestibility can drive higher milk production
Key point → “Corn is corn” is false
Test it → adjust it → model it
Protein Sources Are Not All the Same
“Canola is just canola” → isn't
Protein sources are not created equal
Factors affecting MP supply and efficiency:
RDP/RUP balance
Amino acid profile
Fiber content
Processing method
Canola Meal
Canola meal is a good example of differences between protein sources
Different extraction methods can alter nutrient availability
Studies and meta-analyses show variation in:
Milk yield
N-use efficiency
Depending on the source
Other Protein Sources
The same principle applies to soybean meal and other commodities
Big Picture
Measure DMI → improve feed access → select quality forages → evaluate starch digestibility → evaluate protein sources → adjust the ration → follow the results

Applying Systematic Approaches – Why the Jump Happened
Source + Quality Matter
The proof is simple → source and quality matter
Test your commodities instead of relying on generic values
Use lab values, not generic tags
Goal → protect the MP:ME ratio
Why the Jump Happened
More to Eat → Higher DMI
DMI → dry matter intake
More DMI → more fermentable energy available
MP Matched to ME
MP → metabolizable protein
ME → metabolizable energy
MP matched to ME using tested ingredient values
This allows microbes to run efficiently
Amino acid (AA) supply meets demand
Higher Digestibility
Higher full-tract digestibility of:
Fiber
Starch
More of what is fed becomes nutrients, rather than manure
Outcome
+7 kg milk/cow/day
+0.4 kg fat/cow/day
Achieved by:
Measuring intakes
Testing feeds
Dialling MP:ME using real numbers
Big Picture
Back to basics → but with precision
Measure → test → balance MP:ME → follow results

Technology to Monitor Feed Composition
Feed Composition Monitoring
Technology can be used to monitor the composition of feed resources
These tools help assess the nutritional composition of feeds
Technologies / Systems
ProxiScout
ProxilMate
NIR-Online Solutions
NIRFlex™

Technology in Animal Feeding
Emerging Role of Technology
Technology is playing an emerging role in the feeding of animals
Notably, automation is becoming important in animal feeding
Large Emphasis on “Equipment”
Large emphasis is placed on the “equipment” used for feeding
Automation + equipment are important parts of modern animal feeding systems

Automation in Animal Feeding
Autonomous Self-Driving Vehicles
Autonomous self-driving vehicles are based on:
Sensors
Robotics
A powerful “brain” → needed to process information and make decisions
Automatic Feeding Systems
The same is true for most automatic feeding systems
They require:
Sensors
Robotics
A powerful “brain”
Key Question
Who will develop those systems?

Module 1 – Things We Have Discussed
1. Why Do We Feed Animals?
Why do we feed animals?
2. Important Considerations
What is important to consider when feeding animals?
3. Feed Resources
What do we feed animals?
Various feed resources
4. Feeding Requirements
What determines what and how much we feed a certain type of animal?
5. Food vs Feed
What is the difference between “food” and “feed”?
6. Science + Technology
The role of science and technology in animal nutrition
