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Different components of cognition (including experimental evidence) (look at pic)
Cognition-
The ability of an animal to separate itself from the moment in which it is living contemplate the past predict the future and act accordingly
It involves intelligence, numeric abilities, language, problem solving, etc
Includes all forms of knowing awareness (perceiving, conceiving, imagining, problem solving)
How to measure internal mental states?
Imaging (PET, MRI, etc)
Drawback – interpretation when comparing
Measure behavioral outcomes (observations or direct experimentation)
Theory of mind- and what it allows the animal to do
the capacity to attribute mental states to others
What does this allow an animal to do? -> communication, mating (seeing if a male or female is into you), important to social hierarchies and dominance hierarchy
The mind body problem- the physical mechanical operations of the brain and how does it come to be what we see today
Mirror Self- Recognition Task
If organism can use the mirror to access something on itself it can’t normally see (like look inside of its ear) then they are overcoming the mind body task
Put a mark of red on an animal and see if they are looking and exploring it in the mirror
The elephants are able to pass it
The dot test is done with apes and birds and it works just like in a kid
People debate about what this means
Gaze Following
Deals with understanding knowledge states: if someone is looking over there then maybe it is worth looking there too to see what he is looking at
Did a study with a bunch of diff monkeys and test this sort of thing
In the condition where the test monkey could see that the test monkey was looking at something then they followed his gaze
Could just be classical or operant conditioning at play here
If the animal can use the gaze following to understand knowledge states
Brauer study dominant and subordinate study: had a testing arena when they had treats in the middle of the 2: one treat the dominant couldn't see the other it could: looking at where the subordinate goes-> goes to the one the dominant one can't see shows knowledge states
Santos monkey picks the quiet box deals with attributing knowledge states
Evidence of them using knowledge states
Deception
Can animals be deceptive? What conditions would be necessary for us to consider a behavior actively deceptive?
Yes: doing a behavior the dogs in order to trick the other dogs so that way it can get a treat
active deception in primates
Wheeler (2009) :Put food on a platform: if they find a feeder that is full of food by themseleves then they send out fake alarm calls to get everyone else away
Van Elsakeret al. (2000) :All females outrank men: capuncin monkeys: the males hid the treats from the females and then get them out later when girl gone to eat
le Roux et al. (2015) :When mating both males and females give out calls: when lower ranked mates mate above their rank then they are silent so they don't get yelled at for going above their rank
Empathy
What is empathy and how should we define it?
The ability to project or feel the emotions of another animal
Usually empathy motivates helping
We think the basis for empathy is mimicry
Emotional contagion -> 1 baby starts crying then all the other animals starts crying
Mirror neurons- fire when you perform an action and when you see someone else perform an action
Are learning and memory cognition?
In some ways yes and some ways no
Does it depend on the type of memory?
Semantic- memory for random facts and things (ehh)
Procedural – muscle memory; learning to perform a physical task
Episodic – your personal autobiographically memories (more cognitively different)
Time place learning
The ability to associate a reward or behavioral consequence with a specific location and time, and use that information to return to the location at the specified time in the future
When you feed a pet in the kitchen at a certain time -> they know the time and beg to be fed
Ex the tropical bees and sugar water: they show up 20 min before the time and they stay and drink it; no sugar water then they show up and then quickly leave
Cognitive Maps
A mental representation of an animal's landscape used for calculating optimal routes
Does it demonstrate cognitive ability?
Hard to say: maybe some are making it in a complex map but maybe some are classically conditioned to learn where to go
Problem Solving- Tool Use
Tool use is the exertion of control over a freely manipulable external object (the tool) with the goal of (1) altering the physical properties of another object, substance, surface or medium (the target, which may be the tool user or another organism) via a dynamic mechanical interaction, or (2) mediating the flow of information between the tool user and the environment or other organisms in the environment. (NOT AN EXAM Q)
Evidence of planning ahead in primates for testing like bringing certain rocks to help smash nuts
Counting
Various levels of difficulty
Easy for them to pick between more vs less then
Does seem to be some counting ability
Used a violation of expectation
Animals look at something longer if they find it surprising or interesting
They do a magic show for the animal
Put egg plants behind a curtain and show them putting one behind and either secretly add or take 1 away and the animals stare longer at the things that are shocking (the ones that don't have the correct number of eggplants)
do animals understand probabilities
Yes but with the same cognitive biases that humans have
Some evidence that animals can work in relative terms
Factors influencing animal intelligence
what makes a good test
Intelligence- some combo of quickness to learn, to remember, and solve problems
Good test
Replicability Should get the same results every time you take it
Validity is it actually measuring what you are reporting it to measure
It should have good predictive value it should be able to predict other things well
In animal sense
Must be ecologically valid: must be a task the animal can perform and have a evolutionary relevance to the species
Ex making a giraffe like a predator is stupid
garner’s theory of multiple intelligence
can be bad at 1 but good at something else
Evolutionary Co-option in communication
Evolutionary Co-option and what it happens through
Co-option is the evolutionary adoption of something the animal already does or has for use in a diff form (ex communication)
Happens through
Ritualization – to give something meaning
The intent of the signal must be the same every time
Stereotyping- reduction of variation: boil it down till no variation left
Redundancy- communicating through more than 1 means :Multiple signals trying to get the point across :Reinforces the message helping to eliminate confusion :Ex dogs will urinate and scratch to mark their territory
Communication strategy that has ritualization stereotyping and redundancy
Ex honey bee waggle dance
Different communication modalities (including benefits and drawbacks)
Chemical Signals (Taste and Smell)
Pheromones- chemical signals used in transmitting information WITHIN a species
Carbon based chains
Vary in size and polarity
Small and less polar: likely to be better as airborne signal and break down rapidly : more volatile
Ex sex attractants that moths put out
Larger and more polar: "sticky" more H bonds less volatile/ evaporate slower : stick around longer
Ex good for territory marking
Chemical signals work well in all environments, can linger, can travel far distances
One of the first systems to evolve
What are the advantages and disadvantages?
Advantages
Can be good for a signal to be long lasting
Works well in all environments can work anywhere
Very little chemical noise
Disadvantage
Can be bad to be long lasting -> lead to eavesdropping
Almost no control over its dispersal
Its slow acting
Chemical signals must remain simple by the nature of its chemical structure
Tactile Signals
Touching is efficient; but requires close proximity (speedy but only if you are nearby)
Functionally no "noise" in this system
Can happen in most environments: light or dark
Both affiliative (hi we are friends I'll help you out ex grooming) and aggressive (someone hurting you)
Audible Signals
Sound is vibration through a medium
Energy required to make the vibrations is medium dependent
Drawbacks: Speed is highly medium dependent : Dissipation: further it has to travel the more energy it loses: a drawback to sound: loose its loudness : Can get absorbed by other sources too lowering the intensity
Benefits: work well in any context: not subject to obstacles like being behind someone
Nonlinearity: relationship between pitch and volume is not linear and sort of not 1 to 1 relationship
Body size limits sound amplitude
Largest animals tend to the loudest one
Use low-frequency sounds
Pitch and amplitude
Sound Production – 4 common mechanisms
1. Vibrating a drum – like membrane
2. Stridulating a file and scraper :Think of violin ex in crickets: equipped with a scraper then an adj body part has a file like structure :Both the file and the scraper together –stridulatory organ Pitch altered by how fast you move the structures over one another
3. Vibrating a membrane in air flow :Human vocal chords :Reed in a clarinet
4. Hitting a substrate :Using the environment to create the noise :Ex beavers using their tails to make noise on water
Infrasound
Below what humans can
Orcas and elephants
Things id group identity
Sex age
Ultrasound
Problems that must be overcome to use it
High frequency =short wavelength
High freq =less energy
2 main uses:
Bats, rats, mice, and moths = mating and parenting communication
Echolocation
Advantages using ultrasound
Don't require a lot of energy
Bc of short wavelength much mor efficient of bouncing back
Signals dissipate quickly which is goof for this type of location
Auditory "Noise"
Huge problem in the environment
How can we get around this strategies:
Restrict range of production and perception
Neural ability to sort through both noise and signal
Use in specific contexts
Modern problem: anthropogenic noise (human drived noise)
Visual Signals
Pros: move quickly, cheap to produce
Cons: require light, easily blocked, don't linger
3 main categories:
Pattern on animals surface (including color)
Movement of animals body
Creation of own light source (bioluminescence)
Producing Color Signals
•Pigments- chemicals with c bound structures: either just c or c and n
Longer the chain darker the pigment generally
Pterins (whites, yellows, reds)
Quinones (yellows, reds, oranges)
Malanins(yellowish-brown, brown, black)
Carotenoids (yellows, oranges, reds)
Blue greens are not as common as red oranges and browns
These are almost exclusively seen in birds
There are structure that can create blues as opposed to a pigment
The structure itself absorbs all other colors and reflects back blue
Visual noise- any sort of no intended movement or color
What is a multimodal signal?
A multimodal signal is composed using 2 or more signaling modalities
Ex watching a movie (hearing and sight)
Ex: honey bee waggle dance
Involves visual and auditory cues
3 components to the signal
The angle (tells the degree from the sun)
The amount of time it does this dance indicates the distance
The quality (if it is of good quality it will wiggle harder and faster)
Factors in honest signaling
Honest Signaling
Signals that enhance survival or reproduction will be favored
What happens if a signal isn't honest
Someone could die/get punished: whether it is the false signlaer or the person getting the false signal (they die)
Can lead to an evolutionary arms race
Cheetah and the gazella cheetah gets faster so gazella gets faster and this goes on and on
Recall Theory of Mind
Ex of active deceit in primates: the monkeys gave off false signal of predators so it can hog all of the food
Costly Signals=Honest Signals
The handicap principle states that if only high quality males can exhibit a costly trait and survive, then that trait is an indicator to females of good genes.
Ex peacock tail feather
More necessary when goals of signaler and receiver are not aligned
If goals aligned then there is no reason deceive
Navigation vs. Orientation and the factors necessary to successfully accomplish both
navigation
Navigational- moving with a specific destination
In order to navigate need 2 pieces of info
need to know where you currently are
Need to know how to get where you wanna go
guided movement from 1 location to another but without a specific goal in mind
Navigation is guided movement from one location to another, typically using a compass or landmarks.
To be able to navigate, an organism needs to know at least two things: its current location and the location of its goal. Thus, navigation is not the same as simply moving in a randomly chosen direction, even if that movement is in a straight line
Use environmental clues like landmarks
orientation
Orientation – knowing the direction of where you are and knowing the direction in which you need to go: not goal directed/not super helpful
Orientation is a word that indicates movement in a given direction— classically, movement in a compass direction, although a variety of stimuli that have nothing to do with compass direction may also elicit orientation
Movement in a given direction
Navigation vs orientation ex
If an orienting animal is moving north and it is displaced to the east, it will start from its new location and continue to move due north. If a navigating animal is moving north and is displaced to the east, it will adjust its direction of movement toward the northwest and correct its path, moving to the northwest; in so doing, it will reach the same destination that it had prior to being displaced
Different types of movement
Idiothetic vs allothetic
Idiothetic info- internal to the animal: internally cued info
Ex path integration and homing
Allothetic info: info that is coming externally: environmental cues
Ex: light chemicals smells etc
Direct vs extrapolated search
Direct: using the goal itself as a navigational cue
Depends on things such as communication and interference
Extrapolated: figuring something out within the midst of a diff signals in an environment
Ex when there is a lot of noise during communication you must ignore the noise and focus on the communication
Most common way of locating the source:
Triangulation
Measuring the intensity of a stimulus from 2 diff locations a combo of space and/or time
Simultaneous vs sequential
Ex of simultaneous: the human auditory system
Sequential: doing it all sequential: closing eyes and moving toward the sound and repeating this until you are done
Stereopsis:
the use of 2 simultaneous sensory inputs: ex the 2 eyes of humans taking in info from both sides
Differential importance based off of the life style of the animal
More forward facing the animals eyes are the better the stereopsis they have and better depth perception
Prey animals move eyes to the side of there head so they can have better peripheral vision: allows them to better see predators
A way to componsate a=would just be to constantly swivel head
Kineses v Taxes
Kineses- undirected search: doesn't have the ability to integrate info for multiple sources but can change: ok if simple species in a simple environment
Taxes- changes in directions of movement that are goal oriented with respect to the stimulus source
Orthokinesis-
1 direction movement but not goal directed
Klinokinesis-
turning movement kinesis but not goal directed
Positive taxes-
draws the animal in
Ex positive phototaxis- movement toward light
Negative taxes-
moves the animal away
Ex negative aminotaxis- movement away from the wind
Klinotaxis-
turning that is oriented towards a stimulus
Tropotaxis-
moving towards the average of 2 goals: getting 2 stimuli and moving toward the average of the 2
Telotaxis-
getting 2 sources of info but choosing 1 of them and moving towards it
Ex seeign 2 lights and deciding to move towards the brighter one
Counterturning
When you are forced to move around and object turning in the direction you were previously oriented in
A simple solution for movement around objects
Ex ant in a t maze force him to turn and then he turns in the direction he is trying to go in
Landmarks ( a singular thing)
Using features of the landscape for orientation
Ex tinbergen and the burrowing wasps and how they use landmarks to orient themselves (using pinecone orientations as a landmark)
Name implies a visual landmark but they don't have to be visual: a lot of species use other things
What other cues could be landmarks?
Chemicals or smell
How do animals learn the landscape?
Familiarity, intensity/uniqueness, learning from parents, imprinting
Snapshot Orientation:
The animal remembers the visual scene of the landscape
Also can deal with size: relative size of a landmark I need to move closer or farther
First confirmed with honeybees
Path Integration
The ability to travel to several locations and return directly to point of origin
Simplistic in its mechanics all the animal must do is keep track of the vectors
A lot of animals can do this in the absence of landmarks or cues
Compass Orientation
It appears as if the sun and the moon move around the earth
2 points of reference the north star and south star are constant orientation points
Animals can learn to integrate distances between them
Can be messed up if shift them off the daylight in the artificial environments
Long traveling animals can use magnetic cues to help them determine
Multimodality
- more pieces of info you have the more accurately you can calculate this info
Odometers
Any physiological or mechanical device that allows measurement of distance traveled
Types:
Optic/visual flow- peripheral info that allows you to tell you how fast you are going: speed at which you are going: optic flow ex with pick the shape and color: good for animals with good visual systems
Effort tracker- know how much energy you are outputting and allowing you to tell the distance
Step counting -
Ex study with short legs long legs or normal: either made the ants shorter or taller after they find food source
Hypothesis over shoot or undershoot: animals know how far tongo based on stride length
Cognitive Map
An “internally visualized” landscape
Difficult to distinguish between this, landmarking, and path integration
"that's the only way I know how to get there"
Study with pigeons feed them and then take them to a new place
Fed controls go to rooster site to mate and others go home
Homing-
Repeatedly returning to a central place (nest, territory)
Often after relatively shorter, localized movements
Distinguished from migration
Homed animals are bound
Search may make simple "tools" difficult if not impossible
Have to rely on more complex mind things: path integration etc
Rely on a hierarchy of environmental cues
One of most important pieces of info = distance
Search styles
Random or undirected search-> helpful to start but after a while not very helpful
grid search-> covering a number of turns, used for a shorter more concentrated area
Straight line search -> covers a larger area
Dispersal and migration, including examples and theories explaining them
Migration
Movements of animal populations between seasonally appropriate habitats,
can occur between any two habitats
can occur more frequently than annual (usually winter vs summer but can be more than that)
Can be inherrited genetically, or it can be learned
Differs from dispersal (discussed next) in that return is expected
Also must know the route
Key info: distance & direction
Odometers for distance
And for direction any compass and orientation activities
Bird Migration
Most understood/studied
Twice yearly occurrence –closely linked to circ annual rhythm
Photoperiod
Key factors:
Fat deposition -> store a ton of energy
Restlessness
Both of these things are hormonally regulated
Some species fly nonstop
Stopovers- some need to stop and rest for a bit and fatten themselves up before going on : these places called stopovers
Lot of song birds fly at night even tho active during day why?
To avoid predators
To save energy-> no have to use energy to cool them down
Salmon Migration
Rare fish that changes from freshwater to saltwater and back!
How do they find their way back to natal stream?
Adults first need to find the coast
Cues used: sun geomagnetic, temperature, currents
To find natal stream: must have cues from early life (olfaction)
That is a thing they learn early in life
Trying to smell their way back to the stream they were born in
The Ungulates
vs monarch migration
Purely learned from year to year
From parent to offspring
The Incredible Monarch Migration
Purely genetic: since they no meet previous generation
Dispersal
The process of leaving the natal area and finding a new territory or home range
Climate change can be affecting
Intended to be 1 way
Risky why? -> no guarantee that you can find somewhere just as good as where you are now
One of most morbid periods in an animals life
Who disperses?
Offspring vs parents: Usually it is offspring
Hormonal changes help tell it it is time to leave
Females succeed more than males bc males are happy to accept new mating partners
Honey bees are the opposite
The offspring get to stay and the queen leaves
Males vs females: in mammals males in birds females
Usually in mammals a male must go off and make own herd etc
In birds male hold territory so female must find male who wanna be with
Why Disperse?
Competition :If the pop density too great might be good to go to a new place and get more resources
Avoiding competition :Especially want to avoid competing against own kin so genes are successful
Inbreeding avoidance
Also why 1 sex does the dispersal
Infanticide avoidance :If pop density to high one way to decrease it is to kill off the offspring, so to prevent children from being murdered move away
Forming new colonies
3 Different trophic types and adaptions for their success
Herbivory :Plant eaters
carnivores: meat eaters
Saprophagy :Scavengers/ things that eat dead organic matter
2 types of herbivory and the adv and dis
Browsers
Eat leaves bark and stems from plants
Eat things from higher up/not on the ground
Ex giraffe
Grazers
Eat grass or other things that are at ground level
Ex zebra
Browsers might have a slight adv bc that stuff is around year round even if snow covers grass
But for browsers the areas they are eating are higher in cellulose content and hard to digest and browsing type plants have more plant defense mechanisms
Grazers lower cellulose content and less plant defense mechanisms
Behavioral and gut adaptations
eat fruit when can get it but mostly eat leaves
Leaves high in cellulose
Only leave eaters longer digestive tract and they move slowly
Only fruit eats shorter digestive tract move faster and smarter
Smarter bc more sugar and they must remember where the fruit is leads to more intellectual adaptation
Carnivory
Meat eaters
Sit and Wait Predation
Wait in hiding for a prey to wander by, rather than chasing it down
Advantages:
Conserve a lot of energy
Disadvantage
No idea when what kind and how many prey items wander into your path
How can it increase success?
Sit in an area where the prey areas go so that way they come by
Any lures
Anglur fish that dangles the little light
Cooperative Hunting
Groups of socially bonded animals working together to achieve a goal (obtaining prey)
What conditions are necessary for this to occur?
Need to be able to communicate with each other
Need to know what is going on in the heads of everyone else
Need to live in an area where they can be together
Saprophagy
Scavengers/ things that eat dead organic matter
Feeding on dead or decaying material
Results in a scramble
Advantages:
Don't have to overcome and animals defenses
Don't have to stalk them hunt them chase them down and kill them
Disadvantages:
Usually results in a scramble competition
Whoever is the fastest at eating and digesting it get the most food
Whoever gets there first gets most but then more comp comes
Can result in dangerous situations
Fights between predators
Microscopic competition
Could result in toxic outputs that makes the larger animal sick: might not be good to eat
Optimal Foraging Theory (OFT)
What is the best way for an animal to act
Simple cost benefit analysis
Should be maximizing benefits and minimizing constraints
Benefits- carbs and stuff
Constraints – the energy it takes to get the food
Trying to figure out the point at which you switch behavior
Costs linear
Benefits co linear (like a curve upward)
Trying to model when an animal will continue something or switch
OFT predicts behaviors that maximize fitness
Optimal foraging theory assumes
Natural selection has favored feeding behaviors that maximize fitness
Fitness while feeding increases with energy intake rate
Optimal behavior is the behavior that maximizes fitness
Optimal diet model and what it assumes
The diet model assumes
Foragers maximize fitness by maximizing energy intake rate
Food items are encountered one at a time in proportion to their abundance (can't have multiple food items encountered at once)
Food items can be ranked by their profitability
Profitability = energy /handling time
Handling time= time to manipulate item prior to consumption
Anything greater than 1 Is good you get more energy than you expend
1 you break even
Less than 1 is not worth it you expend more energy than its worth
Average energy intake rate/item = (average energy obtained/item)/ ( avergae search time/item + avergae handling time/item)
oft in crows ex
Methods in crows
Northwestern crows (Corvus caurinus)
Recorded the size of clams that were eaten and the size of clams picked up but not eaten (rejected)
Measured handling times of differently size clams
Measured energy content of differently size clams
Results:
Clams > 30 mm were almost always eaten
Clams < 28 mm were almost always rejected
Seemed like 29 mm was the turning point at that point it seemed to be worth it
Conclusion: Observed crow diets are similar to the diet predicted by the optimal diet model
Marginal Value Theorem
The optimal patch use model assumes:
Foragers attempt to maximize energy intake rate
All patches are identical
Travel time between patches is constant
The instantaneous harvest rate declines as a forager depletes a patch: the forager experiences diminishing returns in each patch
The optimal patch use model predicts the optimal time to spend exploiting each patch
The main thing that determines it is travel time between the patches
A long travel time: stay in the patch you are at
A short travel time: move around a lot
Real life things to consider
Presence of predators -> in the patch and traveling between them
Missed opportunity cost
How much is it costing you to not switch diets etc
Competition with others in the patch