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types of touch senses
tactile, temperature, pain, body sensations
tactile senses are caused by
the mechanical displacement of skin
proprioception
perception of the body mediated by kinesthetic and vestibular receptors
vestibular system tells us the location of our body within the
gravity field
somatosenation
all sensory signals from the body which includes all the components of touch
sensory organ for touch
skin
touch receptors are embedded in the ___ and ____
epidermis and dermis
each touch receptor has these 3 attributes
type of stimulation that the receptor responds to, size of the receptive field, and rate of adaptation (how quickly it fatigues)
4 tactile receptors in the skin
Meissner corpuscles, Merkel cell neurite complexes, Pacinian corpuscles, and Ruffini nerve endings
Meissner receptors are
fast adapting with small receptive fields (FA 1)
Pacinian receptors are
fast adapting with large receptive fields (FA 2)
Merkel receptors are
slow adapting with small receptive fields (SA 1)
Ruffini receptors are
slow adapting with large receptive fields (SA 2)
where is the Merkel cell neurite complex located
at boundary between epidermis and dermis
Merkel cell neurite complexes respond best to
steady downward pressure
Merkel cell neurite complexes important for
texture and pattern perception, fine spatial details
Merkel cell neurite complexes sensitive to
very low frequency of vibrations, < 3Hz
Meissner corpuscles located at
boundary between epidermis and dermis
Meissner corpuscles sensitive to
low frequency vibrations between 3 and 40 Hz
Meissner corpuscles important for
detecting slip of objects across skin to properly grip objects
Ruffini endings are located
deeply in the dermis
Ruffini endings respond best to
sustained downward pressure and lateral skin stretch
Ruffini endings are important for
finger position and grasp
Ruffini endings have a ___ sensitivity to vibrations
low
Pacinian Corpuscles located in
subcutaneous tissue
Pacinian Corpuscles respond best to
high frequency vibrations between 40-700 Hz
Pacinian corpuscles are active when
object makes first contact with skin
kinesthetic receptors
play a role in our sense of where the limbs are and what kinds of movements they make
spindles
convey the rate at which muscle fibers change in length
type 1a sensory fibers
wrap around muscles to detect and signal change in muscle stretch
type ___ sensory fibers adapt while type ____ sensory fibers do not
1a, 2
type 2 sensory fibers
interact with the neuromuscular junctions to signal amount of muscle stretch/position of fibers
thermoreceptors
sensory receptors that signal information about changes in skin temperature
2 types of thermoreceptors
warmth fibers and cold fibers
hot is a sensation of ____, not ___
pain, temperature
nociceptors
sensory receptors that transmit information about painful stimulation that potentially causes damage to the skin
2 groups of nociceptors
A-delta fibers and C fibers
A-delta fibers respond to
initial and quick sharp bursts of pain, strong pressure (crushing) and heat
C fibers respond to
throbbing sensation that evolves after initial surge of pain with sustained stimulation
___ fibers have fast transmission to the brain while __ fibers have slower response
A-delta, C
pain responses can be moderated by
anticipation, religious belief, prior experience, watching others respond, excitement
analgesia
decreasing pain sensation during conscious experience
natural pain relieving effects are caused by
endogenous opiates (block uptake of neurotransmitters that are sending pain sensations to the brain)
gate control theory
a system that transmits pain that incorporates modulating signals from the brain (top down)
gate control theory includes a feedback circuit that is located in the
substantia gelatinosa of the dorsal horn of the spinal cord
pain sensitization
damage to body tissue leaves site being more sensitive to pain
hyperalgesia
an increase in pain sensation
neuropathic pain is pain as a result of
damage to or dysfunction of nervous system
part of the brain corresponding to more cognitive aspects of painful experiences and signalling something is wrong
anterior cingulate cortex
emotional response to prolonged suffering is associated with the
prefrontal cortex
social touch is mediated by
unmyelinated peripheral C fibers aka C tactile afferents
social touch is processed in the
orbitofrontal cortex rather than the somatosensory cortex
C tactile afferents respond best to
slowly moving, lightly applied forces (ex. petting)
touch sensations travel as far as ____ to get from skin of feet to the brain
2 meters
the first synapse for touch sensation occurs at the
spinal cord
first connection occurs at the _____ of the spinal cord
dorsal horn
2 major pathways for touch information once in the spinal cord
spinothalamic, dorsal-column-medial-lemniscal
which touch pathway is slower
spinothalamic
what is the spinothalamic pathway for
pain, temperature, crude touch
what is the dorsal-column-medial-lemniscal pathway for
tactile and proprioceptive information
which pathway has fewer synapses
dorsal-column-medial-lemniscal
spinothalamic pathway
dorsal horn → lateral spinothalamic tract → ventral posterior nucleus of thalamus → cerebral cortex
spinothalamic pathway also provides mechanisms for
pain inhibition
dorsal-column-medial-lemniscal pathway
dorsal horn → Gracile nucleus → medial lemniscus →ventral posterior nucleus of thalamus → somatosensory cortex
primary somatosensory cortex is located on the
postcentral sulcus
how are touch sensations represented
somatotopically, meaning adjacent areas on skin connect to adjacent areas on the brain
phantom limb
perceived sensation from a physically amputated limb of the body
phantom limbs can be felt in areas that are
adjacent to the areas of the amputated limb on the homunculus
2 point threshold
minimum distance at which 2 simultaneous touches are perceptible as separate
typical resolution for temporal details of 2 different touches is
5 ms
auditory temporal resolution is
0.01 ms
haptic perception
knowledge of the world derived from sensory receptors using active exploration/interaction with the environment
exploratory procedure
typical hand movement pattern used to touch objects in order to perceive their properties
lateral motion used to determine
texture
pressure used to determine
hardness
static contact used to determine
temperature
unsupported holding used to determine
weight
enclosure used to determine
global shape/volume
contour following used to determine
global shape/exact shape
fingerprint ridges selectively amplify frequencies from ___ to _____ by a factor of
200-300, 100
ridged fingerprints may help maintain
grip in the presence of moisture
Jack Loomis experiment showed that
accuracy of touch when feeling a raised stimulus is similar to blurry vision
what happens to tactile acuity for sighted people
it gets worse with age
tactile agnosia
inability to identify objects by touch
tactile agnosia is caused by
lesion to parietal lobe
where system of touch
knowing where objects are in the environment when only using touch perception
frame of reference in the where system of touch
coordinate system used to define locations in space
egocenter in the where system of touch
center of a reference frame used to represent locations relative to the body
is there a consistent location of the egocenter?
no
body image
impression of our body in space
tadoma method
method of touch for speech perception for deaf and blind people
physical stimulus for olfaction
odorant
odorant
any specific aromatic chemical that is volatile (able to float through air), small, and hydrophobic
primary purpose of nose
filter, warm, and humidify air we breathe
3 parts of the nose
small ridges (turbinates), olfactory clef, and olfactory epithelium
olfactory cleft
narrow space at the back of the noise where air flows in, where the main olfactory epithelium is located
olfactory epithelium
secretory mucosa whose primary function is to detect odorants
3 types of cells in the olfactory epithelium
supporting, basal, olfactory sensory neurons
supporting cells
provide metabolic and physical support for olfactory sensory neurons
basal cells
precursor cells to olfactory sensory neurons