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Unikonta
Supergroup that includes fungi, animals and some protists (Two Clades: Amoebozoa and Opisthokonta)
Amoebozoa
Clade, consists of true amobas
Opisthokonta
Clade that includes fungi, animals, and related protists
Fungi Synapomorphies
- Feed through absorptive nutrition (decomposers)
- Cell walls with chitin
- Mostly multicellular, some unicellular (yeast)
- Most are saprobes
Absorptive Nutrition in Fungi
They secrete digestive enzymes to break down food, then absorb digested food into their cells
Saprobes
Organisms that live off dead or decaying organic material (Ex: Fungi)
Ecological Importance of Fungi
- Decomposers
- Mutualistic Symbionts (microrrhizae)
- Parasites/pathogens
- Various economic uses
Mycorrhizae
Fungus that grows in association with the roots of a plant, provide water and nutrient absorption (symbiotic)
Endophytes
Harmless fungi inside plant parts, they create toxins to deter herbivores and defend against pathogens
Haustoria
The appendages of parasitic fungi
Hyphae
Filamentous structures that compose a fungus
Mycellium
Mass of hyphae that form the body of the fungus
Septate Hyphae
Hyphae that are divided
Coenoncyte Hyphae
Hyphae that are undivided
Asexual Reproduction in Fungi
Mycelium produces spores that are released, germinate, and form new mycelia
Sexual Reproduction in Fungi
1. Plasmogamy
2. Heterokaryotic
3. Karyogamy
Plasmogamy
fusion of cytoplasm between two mycelia
Heterokaryotic
stage containing two or more haploid nuclei following plasmogamy
Karyogamy
fusion of nuclei to form diploid zygote
5 Clades of Fungi
1. Chytridomycota
2. Zygomycota
3. Glomeromycota
4. Ascomycota
5. Basidomycota
Chytridomycota
Possess flagellated spores, mostly aquatic saprobes or parasites
Zygomycota
- Sporangia in asexual fungi
-Zygosporangia in sexual fungi
-Coenocytic hyphae
- Some form mycorrhizae
Glameromycota
- Mostly endophytes
- Arbuscular mycorrhizae that penetrate root cells
Ascomycota
- Sac fungi and yeast
- Ascus (sac that contains ascospores)
- Ascocarp
- Conidia
Ascocarp
Fruiting body that creates and releases ascospores during the sexual stage of ascomycota
Lichens
- Fungi with a photosynthetic symbiont
- Green algae or cyanobacteria
- Sensitive to air pollution
- Most are ascomycetes
Basidomycota
- Includes mushrooms, shelf fungi, puffballs, jelly fungi
- Basidium (sexual structures)
- Decomposers, pathogens, mycorrhizae and mushrooms
Basidium
- Site of meiosis in basidomycota
- Basidiospores extruded to the outside from the basidiocarp
Archaeplastida
Includes red algae (protists), green algae (chlorophytes, charophyceans), and land plants (embryophytes)
Green Algae
- Chlorophytes & charophytes
- contribute to biofuels
- Both unicellular and multicellular
Chloroplasts
- Stacks (grana) of thylakoids contain chlorophyll a & b
- Store starch
Chlorophyll a & b
Pigments that absorb light for photosynthesis
Terrestrial Adaptations of Plants
- Apical meristems
- Cuticle
- Thick spore walls
- Gametangia
- Dependent embryos
Apical Meristems
Localized regions of plant growth (shoot & stem & root)
Cuticle
Waxy covering to prevent desiccation (drying out)
Thick Spore Walls
Prevent desiccation
Gametangia
Cases that enclose gametes and prevent desiccation
Antheridium
- male reproductive organ of a gametophyte
- produces sperm (haploid = 1n)
- Protects against water loss
Archegonium
- Female reproductive organ of gametophyte
- Produces egg (haploid = 1n)
- protects against water loss
Dependent embryos
Young sporophytes contained within a protective structure
Bryophytes
- Liverworts, mosses and hornworts
- Gametophyte dominant
- Motile (flagellated) Sperm
- No vascular tissue
Hornworts
- Sister taxi of land plants
- Both share stomata
Stomata
Function in gas exchange (CO2 in, O2 out)
Vascular tissue
Duct system for conducting fluids and nutrients; allows for increased height
Tracheophyta
- Sporophyte dominant
- Lignified vascular tissue (xylem & phloem)
- organ systems
Xylem
Conducts most of water and minerals; includes tube-shaped cells called tacheids
Phloem
Cells arranged into tubes to distribute sugars, amino acids and other products of photosynthesis
Lignin
Provide structural support for xylem
Leaves
Organs that increase the surface area of vascular plants and capture more solar energy
Microphylls
Leaves with a single vein
Megaphylls
Leaves with a highly branched vascular system
Sporophylls
Modified leaves with sporangia
Sori
Clusters of sporangia on undersides of sporophylls
Strobli
Cone-like structures formed from groups of sporophylls
Roots
- Organs that anchor vascular plants
- Enable plants to absorb water and nutrients from the soil
- Evolved from subterranean stems
Seedless Vascular Plants
- Formed the first forests
-contributed to global cooling
- Eventually became coal
Seed Plants (Spermatophyta)
- Seeds
- Highly reduced gametophyte
- Heterospory
- Pollen & ovules
Seeds
- Embryo and nutrients surrounded by a protective coat
- Protection, nutrition, dispersion and dormancy
Heterosporty
Evolution of two types of spores (microspores and megaspores)
Microspores
Male spores
Megaspores
Female spores
Pollen
Sperm, derived from microspores
Ovule
Consists of a megasporangium, megaspore, and one or more integuments
Angiosperms
- Flowering plants
- Largest group of land plants
- Flowers, fruits, carpel and double fertilization
Floral Organs
- Sepals
- Petals
- Stamens
- Carpels
Sepals (flowers)
Enclose the flower
Petals (flowers)
Attract pollinators
Stamens (flowers)
Produce pollen (male)
Carpels (flowers)
- Produce ovules (female)
- Ovary at the base and a style leading to a stigma where pollen is received
Filament (flowers)
A stamen with a sac called anther where pollen is produced
Angiosperm Life Cycle
1. Pollen grain germinates in a pollen tube, sperm enters through the micropyle
2. Double fertilization, two sperm enter female gametophyte
3. Fertilization, second sperm develops food storing endosperm
4. Tripoli endosperm nourishes the embryo
5. Double fertilization creates seeds, ovary develops into fruit
4 Fruit Types
- Simple
- Aggregate
- Multiple
-Accessory
Simple Fruit
Develop from a single or several fused carpels (peas, grapes, etc.)
Fruits
Mature ovaries
Aggregate Fruit
Result from a single flower with multiple separate carpels (strawberries, raspberries, etc.)
Multiple Fruit
Develop from a group of flowers (figs, pineapples)
Accessory Fruit
Contain other floral parts in addition to ovaries (pears, apples)
Two Groups of Angiosperms
- Monocots
-Eudicots
Monocots
- Palms, orchids, grasses
- One Cotyledon
- Parallel leaves
- Scattered vascular tissue
- Fibrous root systems
-Pollen grain with one opening
-Floral organs in 3s
Eudicots
- Most angiosperms (roses, daisies, oaks)
- two cotyledons
- netlike veins
- Vascular tissue in rings
- Taproot (main root)
- Pollen grain with 3 openings
- Floral organs in 4s and 5s
3 Organs of Plants
1. Roots
2. Stems
3. Leaves
Root System
- Anchors plant
- Absorbs/stores nutrients
Root Hairs
Where absorption of water and minerals occur; increased surface area
Stem
Organ consisting of alternating systems of nodes (where leaves are attached)
Leaf
- Main photosynthetic organ of most vascular plants
- Flattened blade and petiole (stalk)
Meristems
Perpetually embryonic tissue that allow for indeterminate growth
Apical Meristems
- Elongated shoots and roots through primary growth
- Located near tips of roots and shoots
Later Meristems
In woody plants, increase thickness through secondary growth
Primary Growth
- Produces primary plant body (root and shot system/apical meristems)
Root Cap
Covers root tip and protects apical meristem
Shoot Apical Meristem
Dome-shaped mass of dividing cells at the shoot tip
Leaf Primordial
Where leaves develop along the side of the apical meristem
Auxiliary Buds
Develop from meristematic cells at the bases of the leaf primordia
Vascular Cambrium
- Adds layer of vascular tissue called secondary xylem (wood) and secondary phloem
- one cell layer thick
Pathogen-associated Molecular Patterns (PAMPs)
Molecular sequences that are specific to certain pathogens
Cork Cambrium
replaces the epidermis with periderm (thicker & tougher)
Epidermis
Plant's first line of defense against infection
Effectors
Pathogen-encoded proteins that cripple a plant's immune system
Hypersensitive Response
- Causes cell and tissue death near the infection site
- Induces production of enzymes that attack the pathogen
- Stimulates changes in the cell wall that confine the pathogen
Systematic Acquired Resistance
- Long term response
- systematic expression of defense genes
- Production of salicylic acid triggers defense system to respond quickly