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Plants Architecture
Plant needs
i. collection & conversion of solar energy -> leaves
ii. positioning & support of leaves -> stems
iii. anchorage & absorption -> roots
iv. transport -> vascular system
Leaf Structure
Epidermis, Mesophyll, and Vein (Vascular Bundle)
Epidermis contains what?
- Cuticle
- Guard cells with Stomata
Epidermis
outermost cell layer of a plant body
cutin: (wax) excreted by epidermis
Cuticle
Waxy waterproof covering of a plant.
Produced by the epidermis and has wax to resist desiccation.
Guard Cells
Responsible for opening and closing stomata.
Works together with stomata to regulate gas exchange.
Prevents movement of water ACROSS surface
Stomata
Small openings on the underside of a leaf through which oxygen and carbon dioxide can move.
Mostly on lower surfaces, sometimes on upper surfaces. Found on both surfaces.
Allows for gas exchange.
2 aspects of Photosynthesis
light dependent reactions
light independent reactions
Mesophyll contains what?
- Parenchyma
- Dicots have Palisade & Spongy Layers
Mesophyll
Middle leaf structures photosynthetic layer.
Parenchyma
Ground tissue that forms the bulk of the mesophyll
Can be modified into collenchyma and sclerenchyma
Thin and flexible cells
Most common and versatile ground tissue
Used for metabolic functions and storage of organic products
Palisade & Spongy layers
Palisade layer: where light dependent reactions occur, near the surface. Top part of the mesophyll in dicot plants.
Spongy mesophyll or spongy parenchyma: soft lower layer. Has access to CO2 through stomata. Kelvin Cycle: where carbon fixation occurs, converting nonorganic CO2 into sugars.
Vein (Vascular bundle)
Transports materials throughout the leaf and contains the xylem and phloem.
Separating the VB = damage.
Stem Structure
Epidermis, Cortex, Collenchyma, Sclerenchyma, Vascular Tissues
Xylem
(Xylem Up)
Distributes water from roots throughout
Aimed towards stem
Red
Tells age of tree
Contains: tracheids & vessel members
Tracheids
Thin, hollow, narrow tube, dead cells with perforated, tapered ends.
Vestigial structure
First kind of tube made
Present in early vascular plants and present in angiosperms and gymnosperms (slow flow)
Vessel Members
Thick, hollow, wide tube, dead cells with large holes on end.
Clearly visible
Phloem
(Phloem Down)
Distributes the products of photosynthesis (sugary water) to plants tissues.
Blue
Contains: Sieve tube members (element) & Companion cells
Sieve Tube Members
hollow, living cells with perforated cells
Companion Cells
living cells that help keep sieve tube member cells alive.
Production of sugars in Kelvin Cycle require transportation done by companion cells into phloem
Which 4 cell types is most active metabolically when fully functional?
companion cells
Epidermis in Trees
is replaced by bark or cork. Produced by the cork cambium (tissues that produce other tissues).
Cork (dead) = phloem (alive)
Cork cambium produces cork.
Trees grow wider cause xylem will get clogged
The cork is produced when?
Secondary phloem
Composition of Bark
is produced from phloem, cork cambium, or cork.
Lenticels
cracks in the bark to facilitate gas exchange
Secondary Growth
How a plant increases in girth (diameter)
1. Vascular Cambium
2. Cork Cambium
3. Wood
4. Bark
5. Lenticels
6. "Girdling Plants"
Wood
produced by xylem.
annual rings
Heartwood: clogged xylem, little water transport. Located deeper into trunk, harder wood
Sapwood: newer xylem, free flowing water transport. Phloem makes sap.
Girdling Plants
Weed-Whakers
Cutting a HORIZONTAL band around the circumference of the plant, can be deadly because the vascular cambium, in which nutrients and water travel vertically, can be damaged.
What happens to initial phloem?
it gets crushed
What happens to the xylem?
it gets clogged
Cortex
Yellow layer inside epidermis
Separated by a ring of vascular bundles.
Ground Tissue System
Includes various cells specialized for functions such as storage, photosynthesis, and support
Types of Ground Tissue
Parenchyma, Collenchyma, Sclerenchyma
Collenchyma
celery fibers for support
Sclerenchyma
hard fibers & nodules responsible for support
(ex) rope
Sclerids
nodules glued together to form shells of nuts.
Fibers
secretions that reinforce the stem
Pith
Middle of stem, large because it contains nutrients.
Separated by a ring of vascular bundles.
Replaced by xylem
Procambium
Becomes Vascular Cambium
Makes xylem inward and phloem outward
Grows in rings
Gives rise to vascular tissues
Forms advanced tissues
Monocots
have no cortex
Germ layers
= stem cells
Root Structure
i. Epidermis (permeable) with root hairs
ii. Cortex
iii. Endodermis with Casparian strips
iv. Stele
v. Apoplastic vs Symplastic pathways
Stele
Central cylinder with vascular tissues inside.
Caspian strips
wax, prevents water from growing in between cells
Used to aid a plant and tell weather in the past
Annual rings
Apoplastic vs. Symplastic pathway
Water enters through root epidermis and passes in the spaces "between" cortex cells apoplastically until reaching the endodermis. Casparian strips prevent water from passing between endodermal cells. Thus, water is forced through the cell membranes symplastically where it is filtered before reaching the vascular tissues within the stele = osmosis
Root nodules & Symbiotic bacteria
Bacteria fix nitrogen and are housed in root nodules to supply "fertilizer," thus allowing the plant to thrive, even in soils that are nutrient poor.
Nitrogenace- enzyme that breaks down the triple bond in nitrogen
Microbes contain this enzyme and they're anaerobic.
Mycorrhizae: most plants have an association between their roots and fungi in the soil. This association is critical in aiding water/mineral uptake by the plant.
Abiotic Fixation
Doesn't involve organisms
Lightning converts nitrogen into other sources because N2 isn't a usable form
Less important because lightning doesn't occur enough to be useful for plants
Biotic Fixation
Process by which free nitrogen (N2) is extracted from the atmosphere and converted (fixed) into nitrogen compounds which are plant nutrients (fertilizer). In nature, this process is carried out by certain bacteria such as cyanobacteria
Usable forms of nitrogen for plants
Nitrate NO3
Nitrite NO2
Ammonia NH3
Vegetative Asexual Reproductive modes of flowering plants
Runner (stolon), strawberry
Rhizome- underground stem (bermuda grass)
Corm- modified stem
Tuber- modified underground stem. Gives rise to new growth onion; Potato (however some potatoes are roots)
Bulb- modified stem
Parthenogenesis
development of egg without fertilization
Propagation
vegetative reproductive. Cut off a piece of plant off and it grows.
Plant Development
'... After germination"
Upward growth
-Epicotyl or Coleoptile
-Phototropism
Downward growth
Radicle or Hypocotyl
Gravitropism
Positively Phototropic
Growth in response to light
Structures responsible for downward growth
Radicle and Hypocotyl
Contain statoliths (little rocks) that can sense movement in bottom of cell, triggering the plant to grow towards the center of the earth (down)
Positive Gravitropism
grows in the direction of gravity via statolith sensors
Meristematic Tissues
Plants version of germ cells
Apical Meristems
responsible for increase in plant HEIGHT
Lateral Meristem
responsible for increase in plant DIAMETER (girth)
Meristems vs. Germ Cells
A meristem is the tissue in most plants containing undifferentiated cells (meristematic cells), found in zones of the plant where growth can take place. Meristematic cells give rise to various organs of the plant and keep the plant growing.
A germ cell is any biological cell that gives rise to the gametes of an organism that reproduces sexually.
Meristems are in plants and germ cells are in humans but in essence are essentially preforming the same function
Three Primary Meristems
1. Protoderm = Epidermis
2. Ground Meristem = Parenchyma, Collenchyma, Sclerenchyma = undifferentiated or modified to store pith
3. Procambium = VB w/ xylem and phloem
Exchange & Transport
i. Plants obtain gases, nutrients, minerals, & water via internal fluids
ii. Gas exchange begins with the stomata; roots, lenticils
iii. Internal transport = xylem & phloem
Fluid movement in xylem
Adhesion: Attraction of 2 of different things; water molecules bind to cell wall
This makes sure that the water doesn't go back down (capillary action)
Cohesion: Attraction of two of the same things sticking together; water molecules bind to one another via hydrogen bonding which pulls water molecules upward through xylem, like beads on a string
Evaporation: as water evaporates, it pulls on other water molecules that haven't been evaporated yet
Osmosis: Occurs in roots (root pressure)
Low solute concentration to high solute concentration
Capillary Action: allows water to go up, just a little bit
Transpiration Pull aka Cohesion/Adhesion Tension
The main motive force for transporting water up to the top of a plant (sometimes several hundred feet)
As water evaporates from the leaf's surface the cohesive-adhesive properties of water pull water molecules from below establishing a water tension and pressure
Drawbacks w/ Transpiration Pull
It requires significant water loss from the plant.
In dry conditions or arid environments, this water loss for vertical transport can be critical in plants
Therefore, a replenishing water supply is vital for the roots
Water loss in the tropics doesn't matter cause its always raining there.
Water evaporation = shade
Fluid movement in Phloem
Mass Flow: An active transport mechanism
Source vs. Sink:
Source: Sugars produced by the leaves
Sink: Sugars produced by the rest of the plant. Gravity can assist in this downward movement, however getting the sugars into the cells of the Phloem requires energy
Kingdom Animalia
1. Multicellular
2. Heterotrophs
3. Lack Cell Walls
Two Major Groups of Kingdom Animalia
1. Invertebrates (no backbone) about 15-30 million
2. Vertebrates (chordates with backbones and spinal columns) about 50,000. 30,000 of the 50,000 are fishes
There are animal species that haven't been described yet. 3 million have been described.
Phylum Cortada
vertebrates, fish, amphibians, mammals, birds, reptiles
Metazoa
transition from an animal-like protists to multicellular (choanoflagellate)
Advantages:
1. Large Size
2. Increased Mobility
3. Stable Internal Environment (Homeostasis)
4. Relative independence from environment
Radial Symmetry
Diploblastic
Can be cut in half anywhere along a central axis
No head
Little movement
Bilateral Symmetry
Triploblastic
Can be divided down a central line but cannot be cut anywhere
Cephalization: concentration of sensory structures in the head
(ex) orchids, humans
Asymmetry
No symmetry
Sponges
Ontogeny
= development
"Ontogeny recapitulates phylogeny"
Significant similarities among true appearance of vertebrate embryos due to evolution.
Most abundant skeleton
exoskeleton which tells the shape of the animal
Problem: gets in the way of growth
Largest animal that ever existed on this plant
blue whale
weight (mass) that makes them big
Largest organism on the planet
Humungous fungus
Largest land animal
elephant
Largest invertebrate
squid
First organism to conquer land
Reptiles, by producing a self-enclosed egg (aka shell egg). (ex) birds
# 1 species diversity
tropical rain forests
# 2 species diversity macroscopically
coral reefs
#1 species abundance and diversity
arthropods
#2 species abundance and diversity
mollusks
# 3 species abundancy
Roundworms
1st organism to be cephalized and have excretory system
flatworms
1st organism to have a closed circulatory system and complete digestive tract
ribbon worms
Secondary Compounds
how plants defend themselves from insects, creating medicine.
Almost all animals are
arthropods -> insects -> beetles
Most abundant and diverse
Embryology
i. Cleavage patterns: early cell divisions
ii. Cell fate
Radial Cleavage
new cells placed directly beside or on top of previous cells; deuterosome feature
Spiral Cleavage
new cells placed at juncture between previous cells; protostome feature
Determinate Cell Fate
fate of cells determined early; separated early cells incapable of developing into entire organism; protostome feature
determined @ 4 cell stage
Indeterminate Cell Fate
fate of cells determined relatively late; separated early cells can develop into entire organism; allows for 'twinning' or genetically identical individuals; deuterosome feature
(ex) humans
Developmental Stages
1. Morula
2. Blastula
3. Gastrulation
4. Gastrula
5. Germ Layers
Morula
Solid ball of cells
First stage of development in animals
Blastula
Hollow ball of cells
Blastocoel is in the center
Second stage of development in animals
Blastocoel
Chamber (body cavity) of hollow area within a blastula
Blastocoel is replaced by
mesoderm and coelm