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The properties observed at a particular level arise out of the functioning of parts at the level below → how an ecosystem fucntions is the result of the functioning of the communities within it interacting with the physical environment
To understand the dysfunction in an individual organism, we must look at the the functioning of the organs in that organism. To understand the controls on birth rate in a population, we must look at reproduction in individual organisms
Properties at a given level of organization may be predicted without fully understanding the functioning at lower levels. - predict patterns in ecosystems without understanding all of the details of the dynamics of constituent populations and can predict patterns in populations without understanding all of the details of the responses of individual organism