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clumped dispersion
individuals are aggregated in patches
uniform dispersion
individuals are evenly spaced
random dispersion
position of each individual is independent of each other
territorial
defends space against encroachment by others
demography
study of birth/death/migration characteristics and how they change over time
life tables
summarize survival/reproductive rates of individuals in specific age groups within a population
cohort
group of individuals of the same age
Survivorship curves
plot of percent of cohort still alive at each age
type 1 survivorship curve
low death rates during early/middle life, increased death rate in old age. common in large mammals that produce few offspring but provide them with good care
type 2 survivorship curve
constant death rate over lifespan, ex. rodents, invertebrates, lizards, annual plants
type 3 survivorship curve
high mortality rate among young, later death rate declines. common among organisms that produce large amounts of offspring and provide little care, ex. long lived plants, fish, oysters
R = deltaN / deltaT = B - D
calculates rate of change of population size
dN / dT = rN
represents population growth as rate of change at each instant in time
exponential growth model
only works for ideal environment with unlimited resources
logistical growth model
accounts for the fact that as population increases, resource access decreases
carrying capacity (K)
max population size that a particular environment can sustain
r (K - N) / K
per capita growth rate
rN (K - N) / K
population growth rate
life history
traits that affect organism’s schedule of reproduction/survival
semelparity
individual reproduces once and makes many offspring
iteroparity
individual reproduces multiple times across lifespan
K-selection
selection for traits advantageous in high densities near capacity: nurturing young, etc
r-selection
maximizes intrinsic rate of increase (r ), occurs when density is well below capacity
density dependence
birth/death rate changes based on population density
density independence
birth/death rate does not change with population density
regulated population
1+ density dependent factors cause size to decrease when pop. is large, and size to increase when pop. is negative
resource competition
increasing population density leads to competition between population members for nutrients and toher resources, reducing reproduction rate
metapopulation
when local populations are linked
principle of allocation
individual organisms have a limited amount of resources to invest in functions/activities: there is a trade off
community structure
number of species in community, species that are present, and relative abundance
competitive exclusion
two species competing for the same limiting resource cannot coexist permanently in the same place and eventually the inferior competitor will be eliminated
aposematic coloration
warning bright colors on poisonous species
resource partitioning
differentiation of niches that enable similar species to coexist in a community
fundamental niche
nice potentially occupied by a species
realized niche
portion of fundamental niche actually occupied
character displacement
morphological characteristics diverge more in sympatric species versus allopatric species
cryptic coloration
camoflauge
batesian mimicry
“dishonest”, harmless species mimics appearance of harmful one
Mullerian mimicry
“honest”, 2+ harmful speceis resemble each other so predators learn to avoid them
endoparasites
live inside of hosts
ectoparasites
live outside of hosts
mutualism
(+/+) benefits individuals of both species
commensalism
(+/0) helps one species but is neutral to the other
trophic structure
feeding relationships between organisms
energetic hypothesis
length of food chain is limited by inefficiency of energy transfer along chain
foundation species
have strong effects on communities because of large size or high abundance
keystone species
exert strong control on community structure not by numerical might but by their pivotal ecological roles
ecosystem engineers
species that create / dramatically alter environment
bottom up control
abundance of organisms at each trophic level is limited by nutrient supply/food availability of lower trophic levels
top down
abundance of organisms at each trophic level controlled by abundance of consumers at higher levels. effects move down trophic structure as alternating (+/ -) effects
non equilibrium model
most communities arfe constantly changing after disturbance
climax community
stable equilibrium solely controlled by climate
disturbance
event that changes community by removing organisms or altering resource availability
intermediate disturbance hypothesis
moderate levels of disturbance foster greater species diversity than do high or low levels
small scale disturbances
can create patches of different habitats, maintaining diversity
species diversity
variety of different kinds of organisms in community
relative abundance
proportion that each species represents of all individuals in community
species richness
number of different spedcies in community
Shannon diversity index
-summation (pi ln(pi))
primary succession
process starts on lifeless area. 1st life forms: prokaryotes, protists. 1st macroscopic photosynthesizers: lichenss, mosses from windblown spores. Soil develops gradually and grasses/shrubs can grow
facilitation
when early arrivals in ecological succession make the environment more favorable for later species
secondary succession
recolonization of area after major disturbance removes most organisms in community
species-area curve
the larger the geographic area of community, the more species it has
Island equilibrium model
number of species on an island represents thte balance between immigration of new species and extinction of already present species
intraspecific competition
competition between individuals of the same species
interspecific competition
competition between individiauls of different species
indirect mutualism
if three species are herbivorized by the same herbivore and two of the plants are less palatable, the third plant is eaten more and the other two plants indirectly help each other
Lotka Voterra Predator prey model
negative density dependence enables coexistence, populations fluctuate with each other
spacial grain
characteristic scale where measurements are reported
spacial extent
overall region in which measurements are made at selected spacial grain
abundance
number of individuals
richness
total number of species
evenness
relative similarity in abundance of species
composition
identities of species that are present
alpha diversity
number of species in local site
beta diversity
difference between alpha and gamma diversity
gamma diversity
number of species across all sites
latitudinal diversity gradient
increases in species richness the closer you get to the equator
dispersal
movement of individuals/gametes away from original location
Hadley Cell
air circulation pattern where tropical air heats up, rises, moves away from equator, cools, and moves down
mountain precipitation
precipitation increases at high elevation on windward side of mountains
GPP
total amount of energy from light converted to chemical energy of organic molecules per unit time
NPP
GPP minus energy used by primary producers for cellular respriation
NEP (Net Ecosystem Production)
measure of biomass accumulation by producers and consumers
secondary production
amount of chemical energy in food that consumers convert to their own new biomass
Ecological efficiency
fraction of energy later available to other organisms as growth
assimilation fraction
fraction of energy used by organism for growth/respiration
sociometabolism
metabolism of humans accounting for bodily energy use and indirect consumption through appropriation of ecosystems and other energy sources
flux
rate of movement between compartments
Haber-Bosch process
method of producing nitrogen for fertilizer
Janzen/Connel effect
diseases that increase when hosts have high density population cause negative density dependence for hosts
dI/dt = bSI - mI
rate of infection
R0
bs / m
pyromes
fire dependent biomes
dispersal corridors
pathways that promote persistence of species metapopulation across multiple patches
portfolio effect
higher diversity communities perform better