learning not to respond to a repeated neutral stimulus
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imprinting
learning that can occur only during a specific interval in an animal’s life
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instinctive behavior
an innate response to a simple stimulus
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learned behavior
behavior that is modified by experience
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pheromone
chemical that serves as a communication signal between members of an animal species
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lek
of some birds, a communal mating display area for males
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sexual dimorphism
distinct male and female body forms
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territory
area an animal or group occupies and defends
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dominance hierarchy
social system in which resources and mating opportunities are unequally distributed within a group
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selfish herd
temporary group that forms when individuals cluster to minimize their individual risk of predations
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social animal
animal that lives in a multigenerational group in which members, who are usually relatives, cooperate in some tasks
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altruistic behavior
behavior that benefits others at the expense of the individual
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eusocial animal
animal that lives in a multigenerational group in which many sterile workers cooperate in all tasks essential to the group’s welfare, while a few members of the group produce offspring
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theory of inclusive fitness
alleles associated with altruism can be advantageous if the expense of this behavior to the altruist is outweighed by increases in the reproductive success of relatives
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demographics
statistics that describe a population
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ecology
study of interactions among organisms, and among organisms and their environment
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mark-recapture sampling
method of estimating population size of mobile animals by marking individuals, releasing them, then checking the proportion of marks among individuals recaptured at a later time
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plot sampling
using demographics observed in sample plots to estimate demographics of a population as a whole
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population density
number of individuals per unit area
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population distribution
location of population members relative to one another; clumped, uniformly dispersed, or randomly dispersed
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population size
total number of individuals in a population
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biotic potential
maximum possible growth rate under optimal conditions
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emigration
movement of individuals out of a population
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exponential growth
a population grows by a fixed percentage in successive time intervals; the size of each increase is determined by the current population size
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immigration
movement of individuals into a population
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per capita growth rate (r)
of a population, the change in individuals added over some time interval, divided by the number of individuals in the population
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zero population growth
interval in which births equal deaths
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carrying capacity (K)
maximum number of individuals of a species that a particular environment can sustain; can change over time
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density-dependent limiting factor
factor that limits population growth and has a greater effect in dense populations; for example, competition for a limited resource
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density-independent limiting factor
factor that limits population growth and arises regardless of population size; for example, a flood
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intraspecific competition
competition for resources among members of the same species
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logistic growth
a population grows exponentially at first, then growth slows as population size approaches the environment’s carrying capacity for that species
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cohort
group of individuals born during the same interval
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K-selection
selection that favors traits that allow their bearers to outcompete others for limited resources; occurs when a population is near its environment’s carrying capacity
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life history
a set of traits related to growth, survival, and reproduction such as life span, age-specific mortality, age at first reproduction, and number of breeding events
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r-selection
selection that favors traits that allow their bearers to produce the most offspring the most quickly; occurs when population density is low and resources are abundantsur
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survivorship curve
graph showing how many members of a cohort remain alive over time
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age structure
distribution of population members among various age categories
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replacement fertility rate
number of children a woman must bear to replace herself with one daughter of reproductive age
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reproductive base
of a population, members of the reproductive and pre-reproductive age categories
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total fertility rate
expected number of children a women will bear over the course of a lifetime
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demographic transition model
model describing the changes in human birth and death rates that occur as a region becomes industrialized
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ecological footprint
area of Earth’s surface required to sustainably support a particular level of development and consumption
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commensalism
species interaction that benefits one species and neither helps nor harms the other
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community
all species that live in a particular area
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habitat
type of environment in which a species typically lives
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symbiosis
one species lives in or on another in a commensal mutualistic, or parasitic relationship
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mutualism
species interaction that benefits body species
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character displacement
as a result of competition between two species, the species become less similar in their resource requirements
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competitive exclusion
process whereby two species compete for a limiting resource, and one drives the other to local extinction
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ecological niche
all of a species’ requirements and roles in an ecosystem
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interspecific competition
competition between members of two species
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resource partitioning
evolutionary process whereby species become adapted in different ways to access different portions of a limited resource; allows species with similar needs to coexist
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camouflage
coloration or body form that helps an organism blend in with its surroundings and escape detection
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herbivory
an animal feeds on plants or plant parts
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mimicry
an evolutionary pattern in which one species becomes more similar in appearance to another
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predation
one species captures, kills, and eats another
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biological pest control
use of a pest’s natural enemies to control its population size
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brood parasitism
one egg-laying species benefits by having another raise its offspring
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parasitism
relationship in which one species withdraws nutrients from another species, without immediately killing it
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parasitoid
an insect that lays eggs in another insect, and whose young devour their host from the inside
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indicator species
species whose presence and abundance in a community provides information about conditions in the community
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intermediate disturbance hypothesis
species richness is greatest in communities with moderate levels of disturbance
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pioneer species
species that can colonize a new habitat
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primary succession
a new community becomes established in an area where there was previously no soil
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secondary succession
a new community develops in a site where a community previously existed
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exotic species
a species that evolved in one community and later became established in a different one
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keystone species
a species that has a disproportionately large effect on community structure relative to its abundance
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area effect
larger islands have more species than smalle ones
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distance effect
islands close to a mainland have more species than those farther away
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equilibrium model of island biogeography
model that predicts the number of species on an island based on the island’s area and distance from the mainland
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consumer
organism that obtains energy and carbon by feeding on tissues, wastes, or remains of other organisms
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decomposer
organism that feeds on biological remains and breaks organic material down into its inorganic subunits
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detritivore
consumer that feeds on small bits of organic material
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ecosystem
a biological community and its environment
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food chain
description of who eats whom in one path of energy flow through an ecosystem
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primary production
the rate at which an ecosystem’s producers capture and store energy
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producer
an organism that obtains energy directly from the environment and carbon from inorganic sources; an autotroph
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trophic level
position of an organism in a food chain
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detrital food chain
food chain in which energy is transferred directly from producers to detritivores
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food web
set of cross-connecting food chains
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grazing food chain
food chain in which energy is transferred from producers to grazers (herbivores)
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biogeochemical cycle
a nutrient moves among environmental reservoirs and into and out of food webs
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aquifer
porous rock layer that holds some groundwater
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groundwater
soil water and water in aquifers
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runoff
water that flows over soil into streams
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soil water
water between soil particles
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water cycle
movement of water among Earth’s oceans, atmosphere, and the freshwater reservoirs on land
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atmospheric cycle
biogeochemical cycle in which a gaseous form of an element plays a significant role
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carbon cycle
movement of carbon, mainly between the oceans, atmosphere, and living organisms
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global climate change
a rise in average temperature that is altering climate patterns around the world
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greenhouse effect
warming of Earth’s lower atmosphere and surface as a result of heat trapped by greenhouse gases
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denitrification
conversion of nitrates or nitrites to nitrogen gas
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nitrification
conversion of ammonium to nitrate
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nitrogen cycle
movement of nitrogen among the atmosphere, soil, and water, and into and out of food webs
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nitrogen fixation
conversion of nitrogen has to ammonia
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phosphorus cycle
movement of phosphorus among Earth’s rocks and waters, and into and out of food webs
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sedimentary cycle
biochemical cycle in which the atmosphere plays little role and rocks are the major reservoir
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bioaccumulation
the concentration of a chemical pollutant in the tissues of an organism rises over the course of the organism’s lifetime
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biological magnification
a chemical pollutant becomes increasingly concentrated as it moves up through food chains