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where no community WHAT previously – no WHAT
where no community EXISTED previously – NO SOIL
ex: volcanic island, retreating glacier
begins with WHAT organisms, like lichen (pioneer species) – must be able to survive WHAT conditions
begins with COLONIZING organisms, like lichen (pioneer species) – must be able to survive HARSH conditions – high/low temperatures, extreme dryness, humidity (r-selected species)
develop WHAT to maintain reasonable temperatures and humidity levels
develop MICROCLIMATE to maintain reasonable temperatures and humidity levels
soil building may take from WHAT to a WHAT years
soil building may take from HUNDREDS to a THOUSAND years
What is the order
lichen → mosses→ ferns, shrubs grasses → pines →beeches and maple trees