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N
Estimated number of individuals
M
Number of individuals marked in session 1
n
Number of individuals captured in session 2
m
Number of individuals recaptured in session 2
Lincoln-Petersen Method
N=M(n+1)/m+1
Lincoln-Petersen Method Assumptions
Population is closed
▪ Marking does not affect
probability of survival
▪ Marking does not affect the
recaptures
▪ Marks are not lost, gained, or
overlooked
Edge Effect
Animal home ranges overlap the edge of a sample area, causing population density
estimates to appear higher than reality
Leukemism
Brown in summer, white in winter
True Horns
Unbranched and permanent
• Found only in males or in both
sexes (usually larger in males)
• Grows for animal’s entire life
Bony core: extension of the
frontal bone
Horn: keratinized epidermis
surrounding core of bone.
Antlers
Bony structures that are
branched
o Shed periodically
• Generally males only
Giraffe “Horn”
Pair of short, unbranched, permanent, bony processes known
as ossicones
o Found in giraffes and male okapi
• Situated on sutures between frontal and parietals
• Permanently covered with skin and hair
Rhinoceros “Horn”
Hair-like horny fibers arise from dermal papillae
o Made up of keratin
• Permanent and not branched
• Cemented together to form single horn