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Sounds (Pressure Change)
Created when objects vibrate. This creates pressure changes in the medium.
Longitudinal Wave
a wave in which the particles of the medium vibrate parallel to the direction of wave motion.
Acoustic Reflex
A reflex of the stapedius muscle in the middle ear in response to loud/high intensity sounds to protect the inner ear from potential damage. Follows onset of loud sounds by 200 ms.
Complex Sound
A spectrum that displays how much energy is present in each of the frequencies in the sound.
Fourier Analysis
Break complex waveforms into its pure tone components.
Pitch
The psychological aspect of sound related mainly to the perceived frequency.
Frequency
Is associated with pitch. For sound, the number of times per second that a pattern of pressure change repeats. Low frequency = low pitch. High frequency = high pitch.
Hertz
Measure of frequency. 1 Hz = 1 cycle per second.
Decibel
Unit of measurement for the physical intensity of a sound. Defined by the differences in two sounds as the ratio between the two sound pressures.
Loudness
The psychological aspect of sound related to perceived intensity (amplitude).
Amplitude
The magnitude of displacement of a sound pressure wave. The height of a sound wave. perceived as loudness
Human Frequency Range
20 Hz to 20,000 Hz
Human Decibel Range
0.01 dB to 120 dB
Fundamental Frequency
The lowest frequency component of a complex periodic sound.
Harmonic Spectrum
The spectrum of a complex sound in which energy is at integer multiples of the fundamental frequency.
Timbre
The psychological sensation in which a listener can judge that two sounds with the same loudness and pitch are dissimilar.
Outer Ear
The external sound-gathering portion of the ear, consisting of the pinna and the ear canal. Sounds from environment are collected by the pinnae. Sound waves are funneled by the pinnae into the ear canal. The length and shape of the ear canal enhances certain sound frequencies.
Middle Ear
An air-filled chamber containing the middle bones, or ossicles. The middle ear conveys and amplifies vibration from the tympanic membrane to the oval window. ontains ossicles that transmit sound vibrations from tympanic membrane to inner ear. Uses tensor tympani and stapedius to modulate sound intensity and protect inner ear.
Inner Ear
A hollow cavity in the temporal bone of the skull, and the structures within this cavity: the cochlea and the semicircular canals of the vestibular system canals. Fine changes in sound pressure are transduced into neural signals.
Tonotopic Organization
An arrangement in which neurons that respond to different frequencies are organized anatomically in order of frequency.
Psychoacoustics
The branch of psychophysics that studies the psychological correlations of the physical dimensions of acoustics in order to understand how the auditory system operates.
White Noise
Noise consisting of all audible frequencies in equal amounts.
Tympanic Membrane
The eardrum. A thin sheet of skin at the end of the outer ear canal. Vibrates in response to sound.
Ceruminous Glands
Modified sweat glands, located in external ear canal. Produces earwax.
Malleus
One of the three ossicles. Receives vibrations from the tympanic membrane and is attached to the incus.
Incus
The middle ossicle, connecting the malleus and the stapes.
Stapes
One of the three ossicles. Connected to the incus on one end and the oval window of the cochlea at the other end.
Two Primary Functions of Ossicles
-Transmit sound signals from the tympanic membrane to the inner ear
-Modulate sound intensity before arriving in the inner ear
Organ of Corti
A structure on the basilar membrane of the cochlea that is composed of hair cells and dendrites of auditory nerve fibers. Movements of the cochlear partition are translated into neural signals by structures in the organ of corti.
Hair Cells
Cells that support the stereocilia, which transduce mechanical movement in the cochlea into neural activity sent to the brain stem. Some hair cells also receive input from the brain.
Stereocilia
Hairlike extensions on the tips of hair cells in the cochlea that, when flexed, initiate the release of neurotransmitters.
Cochlea
Spiral structure of the inner ear containing the organ of corti. The cochlea is filled with watery fluids in three parallel canals.
Vestibular Canal
One of the three canals of the cochlea. Extends from oval window at the base of the cochlea to helicotrema at the apex. Canal closest to ossilcles and through which pressures waves move first. Also called scala vestibuli.
Tympanic Canal
One of the three canals of the cochlea. Extends from helicotrema at the apex to the round window at the base of the cochlea. Also called scala tympani.
Middle Canal
One of the three canals of the cochlea. Contains the cochlear partition.
Primary Auditory Cortex (A1)
The first area within the temporal lobes of the brain responsible for processing acoustic organization.
Medial Geniculate Nucleus
The part of the thalamus that relays auditory signals to the temporal cortex and receives input from the auditory complex.
Inferior Colliculus
A midbrain nucleus in the auditory pathway.
Superior Olive
An early brain stem region in the auditory pathways where inputs from both ears converge.
Cochlear Nucleus
The first brain stem nucleus at which afferent auditory nerve fibers synapse.
Auditory Nerve Fiber
A collection of neurons that convey information from the hair cells in the cochlea to the brain.
Conductive Hearing Loss
Hearing loss caused by problems with the ossicles of the middle ear.
Sensorineural Hearing Loss
Most common hearing loss caused by defects in the cochlea or auditory nerve.
Metabolic- Hair cells are injured by chemicals or infection
Sensory- Damage to hair cells by excessive loud noise
Sound Localization
The process by which the location of sound is determined.
Interaural Time Differences (ITD)
The difference in time between sound arriving in one ear versus the other.
Azimuth
The angle of a sound source on the horizontal plane relative to a point in the center of the head between the ears. (what direction is the sound coming from)
0° representing the front (directly ahead).
90° representing the right.
180° representing the back (directly behind).
270° representing the left.
Medial Superior Olive (MSO)
A relay station in the brainstem where inputs from both ears contribute to the detection of Interaural Time Differences (ITDs).
Interaural Level Difference (ILD)
The difference in level (intensity) between a sound arriving at one ear versus the other.
Lateral Superior Olive (LSO)
A relay station in the brain stem where inputs from both ears contribute to the detection of Interaural Level Differences (ILDs).
Cone of Confusion
A region of positions in space where all sounds produce the same ITDs and ILDs.
cues for Auditory Distance Perception
-Relative intensity of sound
-Inverse-square law (as distance from a source increased, intensity decreases faster
-Higher frequencies decrease in energy more than lower frequencies as sound waves travel from a source to one ear
-Relative amounts of direct versus reverberant energy
Absolute Pitch
A rare ability whereby some people are able to very accurately name or produce notes without comparison to other notes.
Vocal tract
Area above the larynx. used for the production of speech. flexibility is important in speech production.
Respiration
Inhalation and exhalation of air. Diaphragm pushes air out of lungs, through the trachea, up to larynx.
Phonation (vocal tract)
The process through which vocal folds are made to vibrate when air is pushed out of the lungs.
Articulation
the act or manner of producing a speech sound using the vocal tract.
The McGurk Effect
an error in perception that occurs when we misperceive sounds because the audio and visual parts of the speech are mismatched.
Vestibular Organs
The set of five organs, three semicircular canals and two otolith organs, located in the inner ear that sense head motion and head orientation with respect to gravity.
Otolith Organs
The mechanical structures in the vestibular system that sense both linear acceleration and gravity.
– Source of our sense of linear velocity and gravity.
Utricle
Otolith organ that has about 30,000 hair cells. Detects horizontal linear acceleration and gravity.
Saccule
Otolith organ that has about 16,000 hair cells. Detects vertical linear acceleration and gravity.
Otoconia
Tiny calcium carbonate stones in the ear that provide inertial mass for the otolith organs, enabling them to sense gravity and linear acceleration.
Semicircular Canals
The three toroidal tubes in the vestibular system that sense angular acceleration, a change in angular velocity.
Anterior, Posterior, Horizontal Semicircular Canals
Filled with a fluid called perilymph. Formed by a membrane filled with fluid called endolymph.
Ampulla
An expansion of each semicircular-canal duct that includes that canal's cupula, crista, and hair cells where transduction occurs.
Vestibulo-ocular Reflex (VOR)
Counter-rotating the eyes to counteract head movements and maintain fixation on a target.
Balance System
The sensory systems, neural processes, and muscles that contribute to postural control.
Active Sensing
Sensing that includes self-generated probing of the environment.
Sensory Reafference
Changes in afference caused by self-generated activity.
Sensory Exafference
Changes in afference caused by external stimuli.
Afferent Signals
Flow from our senses to our brain.
Efferent Commands
Flow from our brain to our muscles.
Graviception
The physiological structures and processes that sense the relative orientation of gravity with respect to the organism.
Spatial Orientation
A sense consisting of three interacting sensory modalities: perception of linear motion, angular motion, and tilt.
Linear Motion
Sensed when accelerating or decelerating in a car.
Angular Motion
Sensed when rotating head from side to side.
Tilt
Sensed as orientation as respect to gravity.
Mechanoreceptor
A sensory receptor that responds to mechanical stimulation (pressure, vibration, or movement).
Push-Pull Symmetry
Hair cells in opposite ears respond in a complementary fashion to each other. When hair cells in the left ear depolarize, those in the analogous structure in the right ear hyperpolarize.
Spatial Cognition
Understanding and organizing 3D space.
Sensory Integration
The process of combining different sensory signals.
Self-Vection
An illusory sense of self motion produced when you are not, in fact, moving.
Proprioception
The ability to tell where one's body is in space.
loudness
The psychological aspect of sound related to perceived
intensity (amplitude)
anterior semicircular canal
Positioned in the vertical plane
-It detects head movements in the sagittal plane, such as nodding the head "yes" (up and down motion).
-It senses rotational movements around the lateral axis (i.e., tilting your head forward or backward).
posterior semicircular canal
It detects head movements in the coronal plane, such as tilting the head from side to side (ear to shoulder).
It senses rotational movements around the sagittal axis (i.e., tilting your head from one shoulder to the other).
horizontal semicircular canal
Positioned in the horizontal plane
It detects rotational movements of the head such as shaking the head "no" (side to side motion).
It senses rotational movements around the vertical axis (i.e., turning your head from left to right or right to left).
study
order:
pinna, ear canal, tympanic membrane, malleus, incus, stapes, oval window, cochlea/vestibular canal, helicotrema, back through cochlear canal, tympanic canal, round window, tectorial membrane, stereocilia, hair cells, spiral ganglion, auditory nerve ochlear nucleus, superior olive, brainstem, inferior colliculus, medial geniculate nucleus, auditory cortex, belt and parabelt area
KNOW: olivary nucleus, inferior colliculus, medial geniculate nucleus
-pinna, tympanic membrane, malleus incus stapes, oval window, cochlea, hair cells, round window, auditory cortex
white noise=white light
-masking other stimuli
ootilith organs: saccule and utricle
know units of frequency (pitch=psychological) and amplitude (loudness=psychological) are measured
analogies:
-hearing=pitch
vision=color
-vision=brightness
hearing=loudness
cues: time and intensity
mcgurk effect: multisensory integration
know how to explain vestibulo-ocular reflex