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when did earth form
4.6 bya
had very little O2
reducing conditions in early earth
reducing conditions favored the synthesis of organic compounds.
to be reducing, atmosphere contains high methane and ammonia
some debate over if atmosphere was actually reducing
first organic compound
may have been near deep sea vents or volcanoes
meteorites may have been another source of organic compounds
synthesis of macromolecules
monomers link to form polymers- process is v slow without enzymes
also faster when concentrated on sand, clay, or rock
protocols
lipids can form vesicles with a lipid bilayer
clay can speed up process
vesicles exhibit simple reproduction + metabolism
self replicating RNAs
first genetic material likely RNA
ribozymes catalyze many reactions in modern cells
RNA may have been a template for DNA
fossil dating methods
Sequence of fossils in sedimentary rock strata
radiometric dating (gives absolute dates)
radiocarbon dating
radiometric dating
isotope half life can be the time required for half the parent isotope to decay→ shows absolute dates
radiocarbon dating
can date older fossils
isotopes with longer half lives cab date tock layers above and below the fossil
Pre Cambian Period
1) 3.5 by a cell life emerged
2) 2.5 bya: oxygen becomes permanent fixture in the atmosphere
3) 1.8 bya eukaryotes orginaite
4) 1.3 bya: multicellur euakatyotes orginicate
oldest known fossil
stromatolites→ rocks formed by the accumulation of sedimentary layers on bacterial mats
oxygen revolution
resulted in prokaryotic groups becoming extinct
survived→ adapt to use aerobic cellular respiration
→ restricted to anaerobic habitats
→ developed tolerance to some O2
ultimate created the ozone layer (O3)→ ozone shield
endosymbiont theory
mitcochonria and plastids were formerly small prokaryotes living within larger host cells
evidence:
- they have their own genomes that are circular
-the organelles transcribe and translate their own DNA
-grow/ replicate outside of the cell cycle
origin of multicellular eukaryotes
1.3 bya
ediacaran biotia (600 myo)→ larver and more diverse soft bodied animals
Paleozoic Era
542-251 mya
Cambrian: sudden appearance of fossils similar to animal phyla
Eras
precambrian, Paleozoic, mesozoic, Cenozoic
periods in Paleozoic era
1. Cambrian: First appearance of many major animal phyla (e.g., arthropods, mollusks).
2. Ordovician: colonization of land by fungi, plants & arthropods
3, Silurian: early vascular plants
4. Devonian: “Age of fishes”; 1 st tetrapods & insects
5. Carboniferous: 1 st seed plants; “forests”; amphibians dominant
6. Permian: radiation of reptiles & insects
Mesozoic Era
(251-65 mya)
Triassic and Jurassic: dinosoucres, gymnosperms are dominant plants
cretaceous: flowering plants Aris
Cenozoic era
65 mya to now
Paleogene
Neogene
Quaternary
-mammals, birds, insects
flowering plants dominant
Historical extinctions/ radiations
boundaries between eras = mass extinction events
permian extinction
250 mya
96% of all marine species
insects and terrestrial life gone
due to high volcanic activity
resulted in large emission of CO2
Cretaceous Extinction
65 mya
50% of Marine species went extinct and terrestrial plants and animals and dinos
partially caused by meterioite impact in Yucatan Peninsula which altered climate and oceanic circulation
Pleistocene Extinction
10,000 years ago
ice ace mammals gone
prob receding of ice sheets
adaptive radiation
evolution of diversity adapted species from a common ancestor
can follow
mass extinctions
evolution of novel charactertics
colonization of new regions
effect of developmental genes
changes In sequence can equal major changed in body form
heterchrony
change in the rate or timing of developmental events
impact body shape
alter reproduction developmental timing
homeotic genes
control placement and organization of body Parts
hox genes: info abt animal development position
prokaryotic traits genome
Genome: prokaryotes have 1 circular chromosome, smaller than eukaryotic
Small rings of DNA → called plasmids
resource use efficiency
small cell size allows more cell surface area per unit volume
allows greater rate of food absorption
can convert food resourced into more biomass via rapid reproduction→ binary fission
nutritional flexibility/ metabolic diversity
ASK
where is cellular respiration is carried out
cellular resp + photosynthesis is in specialized infolding of the plasma membranes
prokaryotic metabolism + O2
obligate aerobes: require O2 for cell resp
obligate anaerobes: NO O2, use fermentation + anaerobic respiration
facultative anaerobes: survive with or without O2
motility
flagella act like boat propellor s
capable of chemotaxis→ moving to/away from a substance
transformation
prokaryotic cell can take up and incorporate DNA from its environment
transduction
movement of genes between bacteria by viruses
conjugation
process where genetic material is transformed from 1 prokaryotic cell to another
donor cell attached to recipient by a plus, then pulls other cell closer to transfer DNA
Archaea Domain
eukaryotes and archaea are more closely related to each other than bacteria
euryarchaeota→ domain archaea
includes halophiles + methanogens
halophiles
require high salinity
methanogens
strict anaerobes→ poisoned by O2
live in herbivore guts, swamps, wastewater treatment plants
crenarchaeota→ domain archaea
includes thermacidophiles→ thrive in hot/ acidic environments
many archaea are not extremophiles
gram positive bacteria
layer of peptidoglycan traps crystal violet
postitive = purple
incredible diversity
ex: staph, strep, actinomycetes
gram negative bacteria
thin layer of peptidoglycan between 2 membranes
doesn’t turn purple
capsule (part of gram positive bacteria)
sticky layer of protein outside cell wall, helps cells adhere to substrate or stick together in groups
endosphere (part of gram positive bacteria)
type of resistant cell that can remain viable in harsh conditions for centuries
cyanobacteria domain bacteria
‘blue green algae”
photosynthetic→ responsible for addition of oxygen to early earth’s atmosphere ‘
chloroplasts likely evolved from this
spirochetes domain bacteria
helical shape
some are free living, others are parasitic
filaments allow for a quince “corkscrew” style of locomotion→ useful to propel through viscous matter
chlamydias domain bacteria
parasites, only live within animal cells
no cell wall, very small'
ex: clamidiya
proteobacteria domain bacteria
gram negative bactiea
e coli
h pylori
mitochronia likely originated from subgroup of proteobacteria
all protests are
eukaryotes
they are in their own kingdom
variety of nutritional modes
photoautotrophs→ make their own food
heterotrophs→ must acquire food from another source
mixotrophs→ combine photosynthesis + heterotrophic
structure of cells
protists are single celled organisms
1 cell must carry out the basic functions
importance of protistan producers
multicellular algae and phytoplankton form the major base of aquatic food webs
protists are primarily
aquatic
protistan diversity stems from
endosymbiosis
primary endosymbiosis + secondary endosymbiosis
excavata
diplomonads, parabasalids, euglenozoans
diplomonads
rely solely on anaerobic pathways
low O2 environments
highly reduced mitochondria
EX: Giardia
Parabasalids
flagellated anaerobes
animal parasites
euglenozoans
presence of spiral crystalline rod inside flagella
euglenids→ commonly found in pond water, photosynthetic
kinetoplastids→ single large mitochondria , some are free living others are parasitic
SAR clade
Stramemophiles, Alveoates, Rhizarians
Stramenophiles
have “hairy” flagellum with smooth flagellum
diatoms, brown algae, oomycetes
diatoms
unicellular algae with glass like cell walls
members of marine + FW phytoplankton
“pillbox” structure→ withstand pressure
brown algae
largest and most complex algae
mulitcelluar→ most are marine
common on temperate coasts with rocky substrate
oomycetes
water molds, white, rusts, downy mildews
tiny filaments that resemble fugal hyphae
distant from fungi
alveolates
dinoflagellates
apicomplexans
cilliates
have alveoli (small membrane- bound sacs) just underneath the plasma membrane
help stabilize the cell and or function in ion balance
dinoflagellates
abundant components of marine and FW phytoplankton
2 flagella located in grooves that run the circumference of the cell
dinoflagellate “blooms” create red tides in coastal waters
some endosymbiotic species live within coral
apicomplexans
animal parasites
have intricate life cycles w sexual and asexual stages requiring different hosts
Plasmodium→ causes malaria
ciliates
use cilia to move and feed
have 2 nuclei (Diff set of genes in each)
normal reproduce asexually via binary fission
eat bacteria or smaller protists
rhizarians
most are amoebas with threadlike pseudopodia
radiolarians
forams
cercozoans
radiolarians
unicellular
planktonic in the ocean
foraminerferans
have a porous shell of CaCO3
live in marine
and FW systems
cercozoans
highly diverse protists
very common in marine, FW, in soil
parasitic or predators-< important for plants/ plant health
archeoplastida
red algae, chlorophytes, charophytes, plants( not a protist)
red algae
red color comes form the pigment phycoerythrin
common in tropical oceans
inhabit very deep waters
green algae
found in aquatic habitats worldwide- also damp soil
chloroplast structure/ pigments→ similar to land plants
have complex life cycles
2 groups: chlorophytes + charophytes
unikonta
have 1 flagellum
amoebozoans + opisthokonts
amoebozoans
different amoebas with lobe or tube shaped pseudopodia
includes: tublinids, slime molds, entamoebas
tubulinids
basic amoebas
common in soil and aquatic environments
active predators
entamoebas
unicellular parasites of animals
slime molds
plasmodial slime molds + cellular slime molds
produce fruiting bodies that aid in dispersal
resemblance to fugi bc convergent evolution
“amoeba” body form is only present during part of their life cycle
plasmodial slime molds
neon colored
decomposers
“acellular” body is 1 giant mass of bytoplams + many nuclei
cellular slime molds
feeding stage consists of solitary amoeba
when food is scare, form and aggregate that functions for reproduction
phytoplankton
unicellular algae, microscopic
seaweed
mulitcellular
different groups have different pigment types
all have chlorophyll A
different accessory pigments
carotenes (orange) → brown seaweed
fleshy (soft) or calcareous (hard) bodies
paper chromatogny
way to separate the combination of pigments
seaweed is not a
plant
seaweed structure
thallus→ entire body
blade→ “leaves”
stipe→ “stem”
holdfast→ holds in bottom but doesn’t function like roots
brown algae
live in temperate regions
intertidal non-kelps
and subtitle kelps
interstitial brown algae
rockweed→ has bumps/ spores on blades
sea palm
pneumatocysts
“airballs” that help seaweed float
subtotal kelp
pacific giant kelp
feather boa
elkhorn kelp
giant kelp
fastest growing organism
forms offshore habitats
economic importance
elkhorn kelp
brown algae 1 large pneumatocyst
long stipe
inertial green algae
sea lecture + dead man’s fingers
tropical green algae
can be calcified
calcareous algae→ contains calcium carbonate
ecological equivalents
similar algae in different geographic areas
intertidal red algae
can be fleshy
filamentous
blades
calcareous
crustose
epiphytic
crustose
grows like a sheet over hard surfaces
resistant to waves + grazing animals
conceptacles
bumps on segments were reproductive spores are shed into the water