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Attenuation
prokaryotes, cis, transcription
transcriptional terminatory secondary structured formed in 5’ UTR of mRNA when trp-loaded tRNAs are available = terminates transcription
riboswitches
both prok and euk (not animals), cis, transcription and translation
Signal binds within the secondary structure changing the shape of the 5’ UTR, can either prevent or cause expression
Transcription- in prok and acts as attenuation
Translation- prok or euk and affects accessibility of ribosome’s binding site (5’ cap in euk and shine dalgarno in prok)
Antisense RNA (asRNA)
in prok and euk, cis and trans, translation, 50-1,000 bp
cis or trans depends on the source of the asRNA
asRNA complimentary to target gene’s mRNA and binds to it and blocks translation by binding to 5’ end covering shine dalgarno in prok or making 5’ end double stranded in euk preventing binding of ribosome
microRNA (miRNA)
euk, trans, translation, 21-25 bp
DICER cuts mRNA into miRNA which binds with RISC which binds miRNA to target mRNA in an imperfect binding format resulting in inhibition
siRNA in mRNA
euk, cis, translation, 21-25 bp
DICER cuts mRNA into siRNA which binds with RISC which binds siRNA to target mRNA in a perfect binding format resulting in degradation
siRNA in DNA
euk, cis, translation, 21-25 bp
DICER cuts mRNA into siRNA which binds with RITS which binds siRNA to target DNA and recruits DNA methyl transferases to methylate the target gene and prevent transcription
long non-coding RNA (lncRNA)
cis, transcription, 200-100000 bp
is transcribed but do not code for a protein, bind to DNA, mRNA, or proteins, can either increase or decrease expression
Ex. Xist in X inactivation
How does Xist work
cis acting, 17,000 pb, lncRNA, expressed from one X csome
RNAs wrap arond X csome that will be inactivated
causes the formation of heterochromatin through histone deacetylation and methylation
inactivated x is known as a barr body
Lyon hypothesis
choice of which x is inactivated is random, solves dosage problem
dosage compensation in drosophila
a set of proteins only expressed in males bind to the x csome and increases x linked gene expression by histone acetylation
dosage compensation in C. elegans
repressor proteins bind to both Xs (two) and repress each about half