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other than subduction, how do rocky shores develop?
glacial retreat
rocky intertidal is stressful
1. physical stress
2. biological stress
3. chemical stress
physical stress
- submersion/exposure
- waves energy
- temperature extremes
biological stress
- space
- competition
- predation
chemical stress
salinity extremes
exposure stress
1. struggle to breath
2. desiccate
wave shock stress
1. Abrasion by particles
2. Hydrostatic pressure from breaking waves
3. Drag from flowing water exerts additional force
controls populations from getting too large
exposure stress gradient
1. spray zone
2. high tide zone
3. middle tide zone
4. low tide zone
how is stress involved in distributions?
puts species in an optimal range due to a balance of abiotic and biotic stress
zonation of rocky intertidal
1. upper intertidal: lichens, periwinkles
2. middle intertidal: barnacles and mussels
3. lower intertidal: seaweeds and surf grasses
conclusions from Connell Experiments
1. Predation is important in lower intertidal
2. Biological factors control lower limit of
species occurrence
3. Physical factors control upper limit
4. Community structure a function of very local processes
how do predators affect community structure?
- sea star, barnacles, and mussels example
- sea stars promote coexistence
- sea stars prefer whatever is most abundant