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Age of life
~3.5-3.8 billion years old
Age of earth
~4.5 billion years old
Age of animals
Animals originated in the Precambrian seas over 600 million years ago
What defines life?
chemical uniqueness, life has a hierarchical organization that does not exist in the nonliving world, reproduction, possession of genetic “program” for inheritance of traits, metabolism, development, movement, evolution, and all living organisms are made up of one or more cells
Chemical uniqueness
macromolecules assembled by living organisms are more complex than those of nonliving matter
general structure of these macromolecules is the same in all forms of life
nucleic acids (DNA and RNA)
carbohydrates
Proteins (composed of amino acids)
lipids
Life has hierarchical organization that does not exist in the nonliving world
atoms
molecules
cells (basic unit of life)
tissues
organs
organisms
populations
communities
Reproduction
living systems can reproduce themselves
Possession of genetic “program” for inheritance of traits
DNA provides the instruction on “how to build proteins”
proteins are necessary for development of unique cells with specialized functions
genetic code translate the sequence of RNA into sequence of amino acids in protein
all forms of life share the same genetic code
Metabolism
living organisms maintain themselves by acquiring nutrients from their environments
metabolism is the sun catabolic (destructive) and anabolic (constructive) rxns in body
Development
all organisms pass through a characteristic life cycle (zygote → adult)
Movement
living systems and their parts show precise and controlled movements arising from within the system
Evolution
Populations of living organisms can evolve over time (i.e., change of allele frequency between generations)
cannot evolve individually, but you can evolve as a population
What are shared traits of animals?
eukaryotes
multicellular
hetertrophs
Eukaryotes
contains membranes-enclosed nuclei and membrane-bound organelles
Multicellular
composed of many cells with specific structure and functions
Hetertrophs
not capable of manufacturing their own food
Evolution (definition)
change of allele frequencies in a population over time
Darwin’s Theory 1
Living world is always changing and changes are passed through heredity
species arise, change and many go extinct
evidence: fossil record
Darwin’s Theory 2
Common descent
all life has descended one ancestor (LUCA)
life’s history can be depicted as a branching tree: phylogeny
more closely-related species share a more recent common ancestor (→ branching) with each other
Evidence for common descent
DNA/RNA, genetic code, transcription and translation, ATP, shared genes
Darwin’s Theory 3
Multiplication of species overtime
every branching point means that one species split into 2
many species go extinct
Darwin’s Theory 4
Gradualism
phenotypically changes occur slowly and gradually eventually producing new species
in contrast: punctuated equilibrium: change happens rapidly and periods of stability can be observed between periods of sudden change
Evidence for both in fossil records
Darwin’s Theory 5
Natural selection (mechanism)
explains how organisms evolved from a common ancestor to become the great variety that can be observed today
natural selection results in traits related to increased survival and reproduction to become more common in a particular environment over time (adaptation)
at the same time, less favorable traits are being lost in the population
Results: natural selection can lead to new traits and species
Homology
similarity attributable to common origin, ie, similarities between (groups of) organisms are explained by a shared common ancestor who passed on the traits
shared traits indicate evolutionary relationship between organisms
Morphological homology
evidence of common ancestry of vertebrates
Developmental homologies
Similarities of embryo development are due to common ancestry
Sequence homology
shared sequence of nucleotides (DNA/RNA) or amino acids (protein) due to shared ancestry
Homoplasy
wings in birds, bats, and insects: used for flying but not due to a recent common ancestor
analogous structures
Natural selection two steps
genetic/phenotypic variations generated as the result oi mutations, genetic recombinations, gene shuffling, etc
sorting (i.e., selecting); survival of different traits
Microevolution
changes in allele frequencies that lead to different traits within a population over time (i.e., pesticides resistance, body size, color, etc)
Macroevolution
changes that lead to new species or taxa over time
What are the 4 evolutionary forces?
selection (natural and sexual)
genetic drift (chance event)
Gene flow (=migration of allele (individuals) out of, into, or between populations)
mutations
Hierarchical organization of animal complexity
protoplasmic
cellular
cell-tissue
tissue-organ
organ-system
Protoplasmic
characteristics of unicellular organisms
complete organisms that can perform all functions of life. Cell organelles are differentiated and perform specialized functions
e.g., amoeba paramecium
Cellular
Multicellular animals have cells combined into larger units
cells are not capable of independent existence cells are differentiated and have specific functions (e.g., reproduction, nutrition, etc) i.e., division of labor
Cell-tissue
aggregation of similar cells into patter of layer that performs a common function (=tissues)
e.g., cnidarians have nerve nets; gastrodermis
Tissue-organ
aggregation of tissues into larger function units, ie., organs
organs are composed of more than one kind of tissue and have more specialized functions than tissue
e.g., heart muscle tissue, connective epithelial and nerve tissue
e.g., platyhelminthes: e.g., eyes, reproductive organs
Organ-system
organs work together to perform a function, i.e., organ system
e.g., circulatory, respiratory, digestive systems
most animal phyla
Lateral
sides
spherical symmertry
any plane through center divides body into mirrored halves
radial symmetry
body can be divided into smaller halves by planes passing through the central axis
Bilateral symmetry
body can be divided into two mirrored parts (L/R) along sagittal plane
medial
midline
ventral
lower (belly) side
dorsal
upper (back) side
Anterior
toward the front
Posterior
toward the back
proximal
describes a body part that is closer to a point of attachment than another body part
distal
describes a body part that is farther away from a point of attachment than another body part
transverse plane
cross section; anterior and posterior
Frontal plane
divides in ventral and dorsal
Sagittal plane
divides in left and right
Acoelom
mesodermal cells completely fill the blastocoel. This cell-filled room; parenchyma
Pseudocoelomate
mesodermal cells lien outer edge of blastocoel; two cavities blastocoel (pseudocoel) and gut
Schizocoel
mesoderm cells fill blastocoel with band of tissue that opens up in center and then forms true coelom (surrounds blastocoel)
4 types of tissues
epithelial
connective
nervous
muscle
3 species concept
morphological (typological) species concept
Biological species concept
Genetic species concept
Morphological (typological) species concept
species are defined by morphological features
Problems: species show variation (e.g., phenotypic plasticity) and do change through time to some degree
Biological species concept
species are groups of interbreeding natural populations that are reproductively isolated from other such groups
Offspring are viable and fertile
Problems: extinct species, asexual reproduction
Genetic species concept
Geneticist’s equivalent of the morphological concept, but genetic similarity is used
problem: how much genetic difference constitutes separate species
8 taxonomic ranks and sequence
domain
kingdom
phylum
class
order
family
genus
species
Ancestral traits
characteristics that was present in common ancestor of all species in taxon
Derived traits
variants that arose later in group
Synampomorphy
derived character shared by all members of clade; evidence of homology and common descent of species in clade
Symplesiomorphy
ancestral, shared character of taxon
Monophyletic
taxon includes most recent common ancestor and all descendants of that ancestor
Paraphyletic
taxon include most recent common ancestor but not all descendants of that ancestor
Polyphyletic
you cannot trace path between all members from group without leaving the group
Endosymbiotic Theory
ancestor of eukaryotes (~2 bya): prokaryote that engulfed another prokaryote
an aerobic prokaryote: today’s chloroplast
a photosynthetic prokaryote: today’s chloroplast
both organelles have their own circular DNA
Common features of unicellular eukaryotes
no germ layers, organs, tissues
mutualistic, commensal, or parasitic (disease) relationship with other species
all requires moist environment
some with endoskeleton, some with exoskeleton, most naked
sexual and/or asexual reproduction
Types of asexual reproduction of unicellular eukaryotes
fission: asexual multiplication
binary or multiple fission
identical cells
Budding: unequal division
progeny cell smaller than parent cell and then grows to adult size
forms of mitosis
Types of sexual reproduction of unicellular eukaryotes
conjugation: exchange of gametic nuclei between paired organisms (e.g., paramecium)
African sleeping sickness
trypanosoma brucei
vector: tsetse fly
Malaria
plasmodium falciparum
vector: mosquito
sticky blood cells block blood vessels (brain, organ)
mosquito → plasmodium in blood stream to human liver: multiple fission (asexual)
then to RBC: feed on hemoglobin (multiple fission)
RBCs burst, releasing parasites and waste products
new RBCs infected
synchronized bouts of bursting and infecting: waves of fever and chills
mosquitoes pick up parasites gametes from host
sexual reproduction: fertilization in gut of mosquito
zygote develops into sporozoite that is transferred to human host
Amoebozoa
Naegleria fowleri
warm water (e.g., ponds, lakesm untreated swimming pools)
usually eats bacteria
enter through nose: water sports
through olfactory nerve to brain
rare infection, usually fatal (~5 days)
symptoms: headache, fever, stiff neck, vomiting, seizures, coma
Evolution of multicellular
singular cells that can do everything
aggregation of undifferentiated cells (e.g., some algae)
cells with different functions and only particular cell types can reproduce (true metazoa)
Who is the sister group of all animals?
choanoflagellates
Choanocytes
generate water stream (move food, waste products oxygen)
Atchaeocytes
ameboid cells that move in mesohyl
can differentiate into other cell: totipotent
Spongocytes
secrete spongin (structural protein in mesohyl)
Sclerocytes
secrete spicules (provide stiffness)
Pinacocytes
flat, thin cells
Types of asexual reproduction of multicellular
Budding: external buds can detach, float away, form new sponges
fragmentation: pieces of sponge detach, float away, adn rebuild sponge (somatic embryogenesis)
Types of sexual reproduction of multicellular
most sponges with sexual reproduction are monoecious
one sponge releases sperm → taken up by another sponge → fertilize egg
zygote → free swimming parenchymula larva → settles somewhere else, adn develops into sponge
What does monecious mean?
Mesohyl
Cells are loosely arranged in gelatinous matrix
Spongin
Spicules
General features of sponges
Important phyla with one example
Movement: whoe does what?, general structure of cilia and flagella, sliding microtubule