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Macrofauna/flora
mammals, most insects, plants, mosses
Mesofauna
nematodes, protozoa
Microorganisms
nematodes, protozoa
Mircroorganisms
fungi, bacteria, actinomycetes, algae
Soil Organism abundance depends on?
quantity & quality of food, environment, predators/competition
Autotrophs
get energy from photosynthesis
Heterotrophs
get energy from consumption of C
Earthworms
segmented worms, burrows, casts; chem/phys properties of soil
Termites
ant-like insects that eat wood/residues; extensive mouths that can alter soil development
Nematodes
unsegmented worms
most prevalent in sandy soils
predators of nematodes, fungi, bacteria, algae, protozoa, insect larvae
plant parasites & plant biological control agents
Protozoa
single-celled, mobile
moist well-drained soils
prey on bacteria
Algae
photosynthesize, some form association with fungi (lichen) that help colonize rock
Fungi
extremely diverse, like acid soils (low pH); includes yeasts, molds, mushrooms
prefer pH 4-5.5
Yeasts
single-celled, often in anaerobic soils
Molds
important in OM breakdown
Mushrooms
high moisture & residue
Mycorrhizae
special type of fungi-plant root association, known to improve P (also Zn) uptake
Actinomycetes
type of bacteria that looks like a fungi
OM degradation (particularly resistant compounds)
produces antibiotics
prefers high pH (7-8.5)
Bacteria
extremely diverse, rapid reproduction
many roles
N reactions
OM & hydrocarbon breakdown
prefer pH 6-7
What are the factors affecting MO growth?
food, environment, predators/competition
What are the effects of organisms on soil?
organic matter decomposition, breakdown of toxic substances, inorganic transformations, N fixation, plant protection/plant pests, physical mixing of soil, competition for O2 when supply is limited
OM Decomposition
C and nutrient mineralization
Residue + O2 = CO2 + humus
termites and other insects involved as well as bacteria, fungi, and actionmycetes
What are the effects of carbon on soil?
soil darkening, CEC, slow release of nutrients, water holding capacity, stabilization of aggregates, infiltration, food for organisms, reduced plasticity of soils (improved manipulation of clayey soils), stabilizes pH, alleviates some toxicities, improves availability of some nutrients
Rate of Degradation
carbohydrates, hemicellulose/cellulose, protein, fats/waxes/oils, lignin, polyphenols/tannins
High C/N Ratio
competition for N = immobilization
Low C/N Ratio
plentiful N = mineralization and plant availability of N
Effects of Lignin on Degradation
reduces immobilization when C/N is high
delays mineralization when C/N is low
Biomass
living
Detritus
dead, but identifiable
Humic Substances
cannot define, but not inorganic
Non-Humic Substances
cannot identify source, but can classify chemically
What are the characteristics of humus?
colloidal, negatively charged, high water holding capacity, dark colored
What happens when you add C?
add residue, compost, manure, green manures
increase plant biomass
How can you reduce losses of C?
reduce harvested biomass
reduce tillage
reduce erosion
Factors affecting Soil C
warm temps, water, vegetation type, soil texture, and poor drainage
cool/wet is HIGH SOM
clay can reduce decomposition and promote fertility
Matric Potential
micropores
Osmotic
water moves toward salt
Gravitational
macropores
Positive soil water potential
moves freely with gravity
0kPa or bars
saturation, water moves primarily via gravity
-10 to -30 kPa (-0.1 to -1/3 bars)
field capacity, water moves primarily via capillary
-1500 kPa (-15 bars)
permanent wilting point, water move primarily via capillary
< 1500 kPa
hygroscopic water, water doesn’t move
Saturated Water Flow
gravitational
Unsaturated Water Flow
capillary
Preferential Water Flow
gravitational