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100 vocabulary flashcards covering Photosynthesis, Systematics, Phylogeny, Taxonomy, Origin of Life, and Course Material from General Biology 2.
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Photosynthesis Summary Equation
The chemical reaction represented by 6CO2+12H2O→C6H12O6+6H2O+6O2, where water is oxidized and light energy drives sugar synthesis.
Thylakoids
Disk-shaped membranous structures arranged in grana inside chloroplasts containing chlorophyll, photosynthetic pigments, and photosystems.
Granum
A stack of disk-shaped thylakoids located within the interior of a chloroplast.
Stroma
The fluid-filled interior space of a chloroplast surrounding the thylakoids where light-independent reactions (Calvin cycle) occur.
Light-dependent reactions
Photosynthetic reactions requiring light that capture energy from sunlight photons, produce ATP from ADP, and reduce NADP+ to NADPH using electrons from water.
Light-independent reactions
Photosynthetic reactions (Calvin cycle) that do not require light and use ATP and NADPH to synthesize organic molecules such as glucose from CO2.
Photon
A particle of light acting as a discrete bundle of energy with no mass.
Photoelectric effect
The removal of an electron from a molecule by light energy, boosting the electron to a higher energy level; discovered by Albert Einstein (1921 Nobel Prize).
Photosystem I
A multiprotein complex in the thylakoid membrane, designated as P700, that absorbs light energy to pass excited electrons to ferredoxin and reduce NADP+.
Photosystem II
A multiprotein complex in the thylakoid membrane, designated as P680, that absorbs light, excites electrons, and uses a water-splitting enzyme to replace lost electrons.
Water-splitting enzyme
Membrane-bound Enzyme 1 inside Photosystem II that oxidizes water molecules to extract electrons, releasing oxygen and protons into the thylakoid space.
b6-f complex
Membrane-bound Enzyme 2 that receives electrons from plastoquinone, transfers them to plastocyanin, and pumps protons from the stroma into the thylakoid space.
NADP reductase
Membrane-bound Enzyme 3 that catalyzes the reduction reaction NADP++H++2e−→NADPH using electrons delivered by ferredoxin.
ATP synthase
Membrane-bound Enzyme 4, a large membrane complex acting as a channel that synthesizes ATP from ADP and Pi as protons diffuse down their gradient back into the stroma.
Plastoquinone
A mobile organic protein electron carrier in the thylakoid membrane that transfers electrons from Photosystem II to the b6-f complex.
Plastocyanin
A mobile organic protein electron carrier that moves electrons from the b6-f complex to Photosystem I.
Ferredoxin
A mobile organic protein carrier that receives excited electrons from Photosystem I and passes them to NADP reductase.
Calvin cycle
The enzymatic pathway of light-independent reactions (also called C3 photosynthesis) in the stroma that fixes CO2 into organic molecules using ATP and NADPH.
Rubisco
Ribulose bisphosphate carboxylase/oxygenase, the enzyme that catalyzes the primary carbon-fixation reaction binding CO2 to RuBP.
RuBP
Ribulose 1,5-bisphosphate, a 5-carbon organic molecule that combines with CO2 during the initial step of the Calvin cycle.
PGA
3-phosphoglycerate, a 3-carbon intermediate compound formed immediately after CO2 is fixed to RuBP by Rubisco.
G3P
Glyceraldehyde 3-phosphate, a 3-carbon sugar produced during the reduction phase of the Calvin cycle that serves as the direct output to form glucose and other sugars.
Photorespiration
An oxidation process where Rubisco binds O2 instead of CO2 to RuBP, favored when stomata close during hot, dry conditions.
Stomata
Microscopic openings on leaf surfaces through which water evaporates and gas exchange (CO2 and O2) takes place.
C3 photosynthesis
Standard photosynthetic pathway where carbon is fixed directly into 3-carbon PGA molecules by Rubisco inside chloroplast stroma.
C4 photosynthesis
A spatial solution to photorespiration where CO2 is initially fixed by PEP carboxylase into a 4-carbon molecule in mesophyll cells before delivery to bundle-sheath cells.
PEP carboxylase
An enzyme in C4 and CAM plants with a high affinity for CO2 that binds atmospheric CO2 to phosphoenolpyruvate (PEP) to form oxaloacetate.
Mesophyll cells
Leaf cells in C4 plants where CO2 is initially fixed into oxaloacetate and converted to malate by PEP carboxylase.
Bundle-sheath cells
Specialized leaf cells surrounding veins in C4 plants where malate is decarboxylated to supply concentrated CO2 to Rubisco for the Calvin cycle.
CAM photosynthesis
Crassulacean Acid Metabolism, a temporal adaptation where stomata open at night to fix CO2 into malic acid stored in vacuoles and close during daylight to run the Calvin cycle.
Aerobic respiration ATP yield
The metabolic oxidation process yielding 38 ATP per glucose molecule using oxygen in the Krebs cycle and oxidative phosphorylation.
Anaerobic respiration ATP yield
Fermentation pathways (alcoholic and lactic acid) yielding only 2 ATP per glucose molecule in the absence of oxygen.
Systematics
The branch of biology dealing with the classification, naming, and evolutionary relationships of organisms based on shared characteristics.
Bacillus anthracis
A Gram-positive, spore-forming bacterium causing anthrax, used in the September 2001 biological mail attacks.
Ames Strain
A highly lethal strain of Bacillus anthracis identified by DNA sequencing during the FBI investigation into the 2001 mailings.
Cladistics
An analytical method of classifying organisms based on shared derived characteristics (synapomorphies) to infer phylogenetic trees.
Derived characteristic
A trait or similarity that evolved in the most recent common ancestor of an entire group and was inherited by its descendants.
Ancestral characteristic
A trait or similarity that arose prior to the most recent common ancestor of a group, exhibited by the outgroup.
Outgroup
An evolutionary reference organism or group that diverged prior to the taxa being studied and possesses ancestral character states.
Synapomorphy
A shared derived character state that originated in the common ancestor of a clade and defines relationships among clade members.
Cladogram
A branching diagram depicting a hypothesized pattern of evolutionary relationships among species based on cladistic analysis.
Clade
A monophyletic group consisting of a common ancestor and all of its descendants, defined by synapomorphies.
Sister species
A pair of species or taxa that share a direct, immediate common ancestor with each other.
Homoplasy
A shared character state between species that was not inherited from a direct common ancestor, caused by convergent evolution or evolutionary reversal.
Convergent evolution
A form of homoplasy where similar traits evolve independently in unrelated lineages adaptively responding to similar selection pressures.
Evolutionary reversal
A form of homoplasy where a lineage re-evolves an ancestral character state, such as the loss of functional limbs in snakes.
Principle of parsimony
An analytical rule favoring the phylogenetic tree hypothesis that requires the fewest evolutionary changes, steps, or assumptions.
Monophyletic group
A taxonomic group that includes the most recent direct common ancestor and all of its descendants.
Paraphyletic group
A taxonomic group that includes the most recent direct common ancestor but excludes some of its descendants (e.g., traditional dinosaurs excluding birds).
Polyphyletic group
A taxonomic group that does not include the most recent direct common ancestor of all member species (e.g., grouping flying vertebrates like bats and hawks).
Binomial nomenclature
A two-part naming system instituted by Carolus Linnaeus consisting of a capitalized genus name and an uncapitalized specific epithet, both italicized.
Taxon
A named group or level of organisms at any tier in the hierarchical classification system (plural: taxa).
Class Mammalia synapomorphies
Diagnostic derived features defining mammals, specifically the presence of mammary glands and hair.
Order Carnivora synapomorphies
Diagnostic derived features defining carnivores, specifically the presence of carnassial teeth.
Family Felidae synapomorphies
Diagnostic derived features defining cats, including a skull with a short rostrum, short facial area, and short inflated braincase.
Genus Felis
A genus of small felines that cannot roar due to a unique throat structure, comprising six recognized species.
Felis margarita
A species of small wild cat inhabiting deserts of northern Africa and southwest/central Asia, characterized by large ear pinnae and reduced size.
Homologous structures
Structures derived from the same ancestral source that may perform different functions, such as dolphin flippers and bird wings.
Analogous structures
Structures performing similar functions that derived from different ancestral sources, such as vertebrate lungs (swim bladder) vs. land snail lungs (mantle tissue).
Sporophyll
A modified leaf that bears sporangia for spore production.
Megasporophyll
A specialized sporophyll that bears ovules, such as those covered in brown ramental hairs in Cycas.

Carpel
The female reproductive organ of a flower, formed by the evolutionary folding of a sporophyll, enclosing ovary, style, and stigma.
Cone scale
Reduced, shortened sporophylls that form the reproductive structures (cones) of gymnosperms.
Hadean Eon
The geologic eon (4,500 MYA−4,000 MYA) marked by the formation of Earth as a hot mass of molten rock.
Archean Eon
The geologic eon (4,000 MYA−2,500 MYA) during which chemically-rich oceans formed and life originated approximately 3.5 billion years ago.
Proterozoic Eon
The geologic eon (2,500 MYA−600 MYA) during which atmospheric oxygen accumulated and the first multicellular organisms appeared.
Phanerozoic Eon
The current geologic eon (600 MYA−Present) marked by abundant visible animal and plant life beginning with the Cambrian explosion.
Cambrian explosion
A dramatic diversification event approximately 500 million years ago in aquatic environments establishing major invertebrate phyla and primitive chordates.
Atmospheric oxygen concentration
The current percentage of oxygen gas in Earth's atmosphere, which is approximately 21%.
Oxygen lag hypothesis
The 200-million-year delay between the origin of cyanobacteria and atmospheric oxygen accumulation, explained by oxygen reacting with iron in oceans to form iron oxide.
Ozone layer (O3)
An atmospheric layer created when O2 interacts with UV radiation, protecting life on Earth by reducing UV-induced mutation rates.
Endosymbiosis hypothesis for mitochondria
Theory stating that mitochondria evolved when a primitive eukaryote engulfed an aerobic Alphaproteobacteria (like Rickettsia or purple non-sulfur bacteria).
Endosymbiosis hypothesis for chloroplasts
Theory stating that chloroplasts evolved when an eukaryotic cell engulfed a photosynthetic cyanobacterium, leading to red and green algae.
Secondary endosymbiosis
The process wherein a eukaryotic cell engulfed a eukaryotic red alga, resulting in brown algae chloroplasts surrounded by four membranes.
Three Domains of Life
The top-level monophyletic clades into which all life evolved: Eubacteria, Archaea, and Eukaryotes.
Dr. Ortega's course extra credit policy
Grants 4 extra credit points per exam for reflection surveys, up to 15 points on the second lowest exam for 100% clicker participation, and 6 points for the Gateway survey.
Exam score dropping policy
The single lowest grade among the four course exams (including the final exam) is dropped when calculating the final exam average.
Weekly quiz weight
Weekly quizzes account for 5% of the final grade, with the three lowest quiz scores dropped.
Exam grade rounding rule
Final calculated numeric course grades are rounded up by adding 0.6 points.
Cell phone & device exam rule
During exams, cell phones must be disconnected inside bags, and smartwatches, earphones, calculators, food, and drinks are strictly prohibited.
Course email requirement
Students must communicate with Dr. Ortega exclusively using their official FIU email address or through the Canvas portal.
Inhalational anthrax mechanism
Inhaled Bacillus anthracis endospores germinate and release toxins in lymphatic system nodes and alveoli, circulating eventually to the brain.
Bruce E. Ivins
A biodefense researcher at Fort Detrick, Maryland who was the primary suspect in the 2001 anthrax mailings before committing suicide in 2008.
Oldest Homo sapiens fossil
The oldest known fossil of Homo sapiens, discovered in Morocco and dated to approximately 300,000 years old.
Ape clade synapomorphy
The shared derived character state defining the common ancestor of all apes, which is tail loss.
Bipedalism
The shared derived character state (synapomorphy) defining the genus Homo relative to knuckle-walking apes.
Cetacean flipper origin
A modified mammalian limb adapted for swimming through convergent evolution, sharing structural homology with bird wings.
Snake limb loss reversal
An evolutionary reversal wherein snakes lost functional limbs possessed by tetrapod ancestors, forming a synapomorphy defining snakes.
Archosauria clade
A monophyletic group containing crocodilians, pterosaurs, dinosaurs, and their direct descendants (birds).
Traditional Dinosaur group classification
A paraphyletic grouping because it historically excludes birds, which are direct descendants of dinosaur ancestors.
Flying vertebrates grouping
A polyphyletic assemblage combining bats (mammals) and hawks (birds) that does not share a single direct flying common ancestor.
Euphorbia handiensis and Cactaceae convergence
Plant taxa exhibiting homoplasy via convergent evolution for CAM physiology, succulent stems, and modified leaf spines in separate deserts.
Land snail colonization
At least 10 independent land colonization events from marine environments, representing convergent evolution of lungs in terrestrial snails.
Terrestrial arthropod colonization
Four independent land colonization events occurring around 400 million years ago by spiders, millipedes/centipedes, crustaceans, and insects.
Aquatic ancestor of insects
The ancestral aquatic lineage of insects identified through phylogenetic analysis as crustaceans.
Alphaproteobacteria
The bacterial group (including Rickettsia and purple non-sulfur bacteria) that gave rise to mitochondria via endosymbiosis.
Cyanobacteria
Photosynthetic prokaryotes originating ∼3.7 billion years ago that generated atmospheric oxygen and became chloroplasts via endosymbiosis.
Mitochondrial DNA
Circular DNA contained inside mitochondria whose nucleotide sequence closely matches Alphaproteobacteria.
Lichen
A symbiotic organism resulting from the intimate interaction between two distinct phylogenetic lineages (a fungus and a photosynthetic partner).
Respiration gene localization
In humans, most of the genes that control cellular respiration are located in the cell nucleus rather than in the mitochondria.