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Nuclear envelope
separates transcription and translation which allows alternative splicing and other forms of RNA processing, controls gene expression; the synapomorphy that defines the domain of Eukarya
Eukaryotic cells are…
larger than prokaryotes, have organelles, and a cytoskeleton
Multicellularity
being made up of multiple cells, evolved multiple times in eukaryotes
Protists
eukaryotes that are not fungi, green plants, or animals; a collection of lineages that are surrounded by water most of the time
Important evolutionary innovations in protists…
a nuclear envelope/ membrane, the mitochondria, the chloroplast
Endosymbiotic theory of mitochondria
proposed by Lynn Margulis; mitochondria evolved from an aerobic bacterium that was engulfed by an anaerobic eukaryotic cell; mutually beneficial- host supplied protection and carbon while bacterium produced more ATP for the host
Mitochondria
similar genes to alpha-proteobacteria, divide independently of the host via fission, have their own ribosomes, have a double membrane, have their own chromosomes
Endosymbiotic theory of chloroplasts
in all photosynthetic protists, originated when a protist engulfed a cyanobacterium, provides host with oxygen and glucose while host provided protection and light
Chloroplasts
similar genes to alpha-proteobacteria, circular genes like in cyanobacteria
Secondary endosymbiosis
in excavata, rhizaria, alveolata, and stramenopila the chloroplast is surrounded by more than 2 membranes (usually 4), when an organism engulds a photosynthetic eukaryotic cell and retains the chloroplasts as intracellular symbionts
Land plants evolved from _______ _______ that lived in freshwater
green algae
Land plants
live in water for water/ minerals, support, and reproduction; are monophyletic; evolved to resist drying out, absorb nutrients, stand upright, and reproduce without water
Embryo protection
nourish a multicellular embryo in the body of the female plant
Cuticle and stomata
cuticle is a waxy sealant that prevents water loss and inhibits the uptake of CO2, stomata are pores bounded by guard cells that allow the uptake of CO2 and control water loss, gas exchange done through pores
Non-vascular plants (Bryophytes)
simplest plants, lack roots and vascular tissue, water and nutrients move by diffusion, small, depend on water for reproduction; ex: mosses, liverworts, and hornworts
Roots
absorb water and key nutrients; made up of taproots and lateral roots; anchor plants, absorb ions and water, obtain energy from sugar, store material
Shoots
absorb light
Vascular tissue
what distinguishes vascular and non-vascular plants; made up of xylem and phloem; have tracheids to conduct water; increases structural support
Xylem
transports water and mineral from roots upward
Phloem
transports sugar, amino acids, hormones, and other substances from root to shoots and shoots to root; made up of sieve-tube elements to connect cytoplasm of cells and companion cells to support metabolic function
Wood
formed in stems and branches of some vascular plants, tracheids and vessels come together
Transpiration
water transport from roots to shoots due to different water potential; driven by evaporation of water from stomata of leaves
Conditions for transpiration
stomata are open, the air surrounding leaves is drier than the air inside of leaves
Cohesion-tension theory
open stomata create a water potential gradient between the interior of a leaf and its surroundings, water evaporates from the cell wall of plants, a negative force pulls water from root to leaf
Translocation
the movement of sugars through the plant by bulk flow from sources to sinks through the phloem
Sources
the tissue where sugar enters the phloem, high sugar concentration
Sinks
tissue where sugar exits the phloem, sugar concentration is low
Pressure-flow hypothesis
explains how sugars are moved through the phloem, water in phloem sap moves down a pressure gradient carrying sugars; high turgor pressure near source and low turgor pressure near the sink; moves sap toward sinks
Seedless vascular plants
first to evolve vascular tissue to move water and nutrients upward and dissolved sugar down leaves, use spores and the wind for reproduction
Pollen grain
in land plants, allows plants living in dry habitats to reproduce efficiently, can be exposed to air for longer periods of time, carried to female plants by wind or animals
Seeds
portable embryos that can disperse to new locations, allows for less competition between plants, distinguishes plants with seeds from seedless plants
Vascular seed plants (Gymnosperms)
first plant to evolve seeds, use seeds rather than spores, have naked seeds many times come in cones, do not need water and do not have flowers and fruit
Flowers
reproductive structures that attract pollinators and develop into fruit, distinguishes angiosperms from gymnosperms
Pollination
the transfer of pollen from one individual stamen to another individuals carpel
Directed-pollination hypothesis
natural selection favored structures that rewards an animal for carrying pollen directly from one flower to another
Flower variations
vary in size, structure, scent, and color in order to attract specific pollinators
Flowers attract pollinators by providing them with _______
food; either protein-rich pollen or nectar or sugar-rich fluid
Trickery
a strategy for achieving pollination, a plant deceives animals into transporting pollen
Bribery
a strategy for achieving pollination, a plant offers something of value to pollinator
Diversity of angiosperms resulted from __________ with animal pollinators
coevolution
Fruit
a structure derived from an ovary and encloses one or more seeds, a fertilized egg becomes a zygote, an embryo plus stored food grows into a fruit, allows for efficient seed dispersal
Animal-dispersed seeds/fruit
fruits are colorful and conspicuous, taste good, usually good for animals
Angiosperms
vascular plants with covered seeds, only plants to have flowers and produce fruit, highly diverse - great adaptive radiation
Adaptive radiation
single lineage produces large number of descendant species that are adapted to wide variety of habitats
Animalia
multicellular eukaryotes that move under their own power, have both nerve cells and muscle cells, ingest their food prior to digestion
Sponges
earliest animals to appear in the fossil record, do not have nerve or muscle cells, part of a paraphyletic group - some but not all descendants of a common ancestor
Diploblasts
animals whose embryos have two types of tissues or germ layers, consist on an ectoderm and an endoderm
Triploblasts
animals whose embryos have three germ layers, consist of the ectoderm, endoderm, and mesoderm
Ectoderm gives rise to ____________ and ____________ _____________ in triploblasts
skin; nervous system
Endoderm gives rise to the lining of the __________ __________ in triploblasts
digestive tract
Mesoderm gives rise to ____________ ____________, ___________, and _____________ ___________ in triploblasts
circulatory system; muscle; internal structures
Cilia
used by ctenophores to swim
ctenophores and cnidarians lack true ______________ ____________
mesodermal muscle
Contractile proteins
all animals share homologous genes for this, the proteins responsible for muscle
Epitheliomuscular cells
proteins in ctenophores and cnidarians used in contractile cells, similar to true mesodermal muscle cells but evolved independently
Radial symmetry
multiple planes divide the organism into mirror images
Bilateral symmetry
only one plane divides the organism into mirror images
Bilaterians
have diverse central nervous systems, tend to encounter their environments at one end
Cephalization
the evolution of a head where structures for feeding, sensing environment, and processing information are concentrated, where the brain is located and coincided with the evolution of the CNS
“Protostome”
develops mouth first, the gut develops from front to back
“Deuterostome”
mouth forms second, the gut develops from back to front
Cambrian explosion
marine animals evolved most of the basic body forms, lots of fossil evidence of relatives of todays species
Protostomes
make up atleast 22/ 30 known animal phyla, contains almost all animal species
Bilaterian body shape
inner tube of a gut with a mouth on one end and an anus at the other and an outer tube that forms the nervous system and skin, mesoderm forms muscles and organs
Coelom
an enclosed, fluid filled body cavity between the tubes in a bilaterian, provides space for oxygen and nutrients to circulate, enable the internal organs to move independently of each other
True coelomates
bilaterians whose coelom is completely lines with mesoderm
Acoelomates
bilaterians that have no coelom, such as flatworms
Pseudocoelomates
bilaterians whose coelom is only partially lined with mesoderm, like roundworms and rotifers
Molting
shedding an exoskeleton and replacing it with a larger one at regular intervals
Deuterostome
high diversity in bodies, feeding, movement, and reproduction
Echinoderms or spiny-skins
have spines or spikes, marine, very abundant especially in deep waters
Hemichordata
acorn worms and pterobranchs, burrow, deposit-feed or suspension-feed
Chordata
includes lancets, tunicates, vertebrates
Notochord
a rod of tissue from head to tail, stiffens when muscles contract during locomotion, sets chordates apart
Dorsal hollow nerve cord
a bundle of nerve cells that runs from brain to posterior, forms the central nervous system in vertebrates, sets chordates apart
Pharyngeal gill slits
gills, in aquatic chordates, separates type of chordate
Hagfish
have cartilage, lack jaws and vertebrae, have a notochord, swim like snakes, produce lots of slime
Vertebrates
monophyletic group distinguished by synapomorphies: the vertebrae and cranium
Vertebrae
protects the spinal cord, column of cartilaginous or bony structures, forms along the dorsal side of most species
Cranium
protects the brain and sensory organs; bony, cartilaginous, or fibrous case encloses the brain
Vertebrate brain regions
forebrain, midbrain, and hindbrain, key innovation in vertebrate evolution
Lamprey
a vertebrate, cartilage skeleton, jawless, parasitic, suck on other fish fluids
Paired fins
allow the fish to swim fast and straight
Jaws
enabled predators to capture prey and bite chunks
Fish
aquatic vertebrates with gills and fins, most diverse and largest group of vertebrates
Limbs from fins hypothesis
a series of species shows a gradual transition from a lobe-like fin to a limb that could support walking on land, problems included respiration, gravity, and egg desiccation
Amphibians
first terrestrial vertebrates, live in water and on land
Amniota
terrestrial vertebrates, lineage that includes all tetrapod’s other than amphibians, have an amniotic egg, reptiles and birds (lay eggs)
Amniotic egg
a protective covering that reduces the rate of drying
Reptiles
amniotes, have paired kidneys, skin covered in scales, ectothermic
Feathers
evolved before birds, in many species of dinosaurs, not used for flight but rather courtship, territorial displays, and insulation
Mammals
a monophyletic group of amniotes named for mammary glands, produce milk, have cheek muscles and lips, endotherms, have hair or fur
Thermoregulate
control body temperature
Conduction
the direct transfer of heat between two physical bodies that are in contact with each other
Convection
a special case of conduction in which heat is exchanges between a solid and liquid or gas rather than being two solids
Radiation
the transfer of heat between two bodies that are not in direct physical contact
Evaporation
is the phase exchange that occurs when liquid water becomes a gas
Endotherm
an animal that produces adequate heat to warm its own tissue, extremely high basal metabolic rates
Ectotherm
an animal that relies on heat gained from the environment, only generate small amounts of heat as a metabolic byproduct, need less food
Homeotherms
keep their body temperature constant