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Diploid number (2n)
The total number of chromosomes in a diploid cell.
the structure of a DNA molecule
is key to its ability to transmit genes from one generation to the next
local adaptation
when a population of organisms has evolved to be more well-suited to its environment than other members of the same species
Evolution
Change over time
Evolutionary Theory
A concept developed by Charles Darwin to explain the ways in which animals adapt to their environments
natural selection
the process whereby any heritable features that enhance fitness of an organism, relative to its peers, increase in frequency in the population in succeeding generations
fitness
Ability of an organism to survive and reproduce in its environment
Variation
differences in physical traits of an individual from the group to which it belongs
Inheritance
The process in which genetic material is passed from parents to their offspring. (encoded in DNA)
selection
environmental pressure
competition for resources
A cause of conflict that occurs when the demand for resources is greater than the resources available.
Adaptation
A trait that helps an organism survive and reproduce, change in phenotype
Scala Naturae (Great Chain of Being)
The scala Naturae (scale of nature) was a Greek notion for how to rank the biological world. At the bottom were "simple" organisms, like worms, and at the top were " complex" organisms and at the very top of the heap were humans, which were seen as being the most god-like of organisms. This ranking system was borrowed by Christian thinkers where it was expanded into the Great Chain of Being. It's the same ranking system, but here the ranking includes various classes of people, various types of angels, and goes on up to God. I bring this up in class because it speaks to this notion that we often have that sees humans as being apart, or distinct from, the rest of nature. Thus, natural laws are often thought not to apply to humans. This is part of that notion that I spoke of as the "the special place of humans in nature." It's one of the biases, or misconceptions, that people have. According to this principle, humans could not have evolved.
Al-Jahiz
an Islamic scholar who lived in the 800s and who presented a theory about the evolution of animals.
Al-Beruni
from 973-1048 AD, Turkmenistan/Uzbekistan, Muslim evolutionary scholar
- Similar to Ibn Miskawayan, Al-beruni held the view that humans migrated through various "lower" lifeforms on the way towards humanity
- Noting similarity between monkeys and humans, he stated monkeys were the final through which humans passed before coming human
Fixity of species
all species remained unchanged throughout the history of the earth
Carlos Linnaeus
Father of taxonomy, Swedish botanist, zoologist, taxonomies, and physician. He established the system of binomial nomenclature.
binomial nomenclature
Classification system in which each species is assigned a two-part scientific name
Systema Naturae (1735)
Carl von Linnaeus- tried to classify every living things
Linnaean Classification System
Kingdom, Phylum, Class, Order, Family, Genus, Species
Kingdom
First and largest category used to classify organisms
Phylum
Group of closely related classes
Class
in classification, a group of closely related orders
Order
in classification, a group of closely related families
Genus
A group of similar species
Species
A group of similar organisms that can breed and produce fertile offspring.
King Phillip Came Over For Good Spaghetti
Kingdom, Phylum, Class, Order, Family, Genus, Species
Rules of Linnaean Classification
-Unique name
-Latin or Greek
-Genus and species in italics
-Genus is capitalized
-Species is lowercase
-A species without a genus is meaningless
Linnaean Classification of Humans
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Primates
Family: Hominidae
Genus: Homo
Species: sapiens
Buffon (1707-1788)
Noted that different geographical regions had slightly different looking plants and animals despite similar habitats. Environment could change species, but not create new species. Idea that species are not completely static; they may change and the environment may drive that change.
Erasmus Darwin
Also proposed the idea that species could change over time due to environmental changes.
Idea that living species descended from common ancestors.
(So the idea of evolution existed before C. Darwin, but no mechanism to explain how it worked)
Lamarck's theory
an organism acquires its trait through its lifetime and passes it to offspring (use and disuse). We now know that genetics do not work as Lamarck imagined, but his theory was accepted by many for a number of years (one can't inherit most acquired characteristics from one's parents)
George Cuvier
Developed the theory of catastrophism
Catastrophism
A principle that states that geologic change occurs suddenly by sudden events
Charles Lyell
effectively discredited the long-standing view that the earth's surface had been formed by short-lived cataclysms, such as biblical floods and earthquakes-his principle: uniformitarianism: same geological processes that are at work today slowly formed the earth's surface over an immensely long time
Uniformitarianism
Charles Lyell's idea that geologic processes have not changed throughout Earth's history.
Thomas Malthus
Said that populations increase faster than food supplies
Alexander von Humboldt
"father" of modern physical geography/biogeography
Charles Darwin
English natural scientist who formulated a theory of evolution by natural selection (1809-1882)
Voyage of the Beagle
Charles Darwin's famous global voyage, where he found his first evidence of evolution
Galapagos Finches
cactus eater
seed eater
insect eaters
The Galapagos Finches demonstrate natural selection through adaptation.
Origin of Species
1859: Charles Darwin's book explained how various species evolve over time and only those with advantages can survive and reproduce. Darwin built off of concepts from Lyell and Malthus
Darwin's mechanism for evolution
natural selection and genetic variation
adaptive radiation
An evolutionary pattern in which many species evolve from a single ancestral species
Alfred Russel Wallace
British naturalist who developed a hypothesis of natural selection similar to Darwin's
Wallace Line of Biogeography
separates flora and fauna of islands as associated with mainland Asia or mainland Australia
Natural selection (Darwin's)
1) More offspring are produced than the food supply can support
2) There is variation among individuals withinnall species
3) There is variation on heritable traits
Darwin hypothesized that those individuals with variations that allow them to better compete will survive and reproduce more frequently than those individuals without these favorable traits (i.e.., greater fitness). Populations diverge over time to form new species.
Natural selection is not
just survival of the fittest, nor is it the only force of evolution. other foces include mutation, migration, and genetic drift
genetic drift
change in the genetic composition of a population from generation to generation ti the chance sampling events
founder effect
extreme change in the genetic makeup of a population when only new population is established by only a few individuals.
inheritance of acquired characteristics
Lamarckian belief that characteristics acquired during the lifetime of an organism can be passed on to offspring
common ancestry (common ancestor)
a group of organisms share common descent if they have a common ancestor thus all living organisms on Earth are descended from a common ancestor. It explains why different species share similar traits.
adaptations
Changes in physical structure, function, or behavior that allow an organism or species to survive and reproduce in a given environment.
Patterns of Evolution
anagenesis, stasis, cladogenesis
phyletic gradualism
gradual change in the morphology of a population of organisms through time, either anagenetic, cladogenetic, or both (Darwin thought of evolution in this way)
Prokaryotes vs. Eukaryotes
- Prokaryotes have no nucleus or membrane bound organelles
- Eukaryotes have a nucleus and membrane bound organelles
- Both can reproduce and respond to the environment
RNA
ribonucleic acid, a molecule in the cell, which contains genetic information
haploid number (n)
the number of homologous pairs in a diploid cell.
meiosis
occurs to form gametes (sex cells), one cell divides into 2 cells, then those 2 cells divide again into 4 cells.
4 bases of DNA
Adenine, Thymine, Guanine, Cytosine (or A, T, G, C)
DNA bases
weak hydrogen bonds. each base only binds with a certain other base, each pair known as a "base pair"
DNA replication
the process of making a copy of DNA
DNA transcription
the formation of an RNA strand complementary to the DNA strand by RNA polymerase
gene
region of DNA that codes for a specific protein, a basic unit of heredity in a living organism
Transcription
DNA sequence in a gene is copied into mRNA
Translation
RNA is translated into an amino acid chain (or protein)
for transcription, thymine (t) basss are replaced by what as the complementary base to adenine?
uracil (u)
amino acids
building blocks of proteins
alternative splicing
splicing can occur more than one way to produce different products (same gene, different products)
protein coding genes
Genes that encode instructions for making proteins.
regulatory genes
genes that control gene expression
Pangenesis
(outdated) Darwin's hypothetical mechanism for heredity- reproductive cell contains gemmules or invisible germs derived from an individual. those eventually get to gametes and pass on traits to their offspring
phenotype
An organism's physical appearance, or visible traits. (observable expression of an organism's genotype)
With genes, we can trace heritability
use laws of probability to compute how alleles will be distributed
Law of Dominance
In many traits one allele is dominant over the other allele. The "weaker (recessive" allele is only expressed when it is paired with another recessive allele
Homozygous
Having two identical alleles for a particular gene (TT or tt)
simple traits
Traits whose variations are controlled by different versions of a single gene (mendel's traits are these) (they include pea color, pea texture, Hitchhiker's thumb, detached ear love, certain diseases)
Complex traits
traits that are influenced by more than one gene (polygenic) or environmental factor (such as height, skin color, many inherited diseases)
population
a group of organisms of the same species that live in a specific area
population genetics
the study of how genes (alleles) evolve in populations
the study of allele frequency distribution and change under the influence of four evolutionary forces
Hardy-Weinberg equilibrium
- a populations allele and genotype frequencies are constant unless there is an evolutionary force acting on them
- equation that was developed to determine the frequency of two alleles at one locus in a population
- mathematical model which we compare null prediction against actual observed data
- if A population is in "Hardy-Weinberg Equilibrium" then evolution has NOT occurred
Mathematics of Hardy-Weinberg Equation
(p^2)+(2pq)+(q^2) = 1
p is the frequency of dominant allele
q is the frequency of the recessive allele
(p^2) is the homozygous dominant
(2pq) is heterozygous
(q^2) is homozygous recessive
in a population with no evolution, genotype ratios will remain constant
natural selection allele frequencies
if one phenotype is associated with higher fitness (greater ability to survive and reproduce), then alleles that result in that phenotype will increase in the population
-evolution by natural selection at the genetic level
sexual selection
A form of natural selection in which individuals with certain inherited characteristics are more likely than other individuals to obtain mates.
mutations
errors which happen during DNA replication
insertion mutation
a mutation in which one or more nucleotides are added to a gene
deletion mutation
a mutation in which one or more pairs of nucleotides are removed from a gene
X-linked disorders
disorder is on the X chromosome and can affect both female and males depending on the carrier. X linked disorders skips a generation- fathers don't pass it on to sons, daughters are carriers and have a 50% chance of getting the disorder because they inherit one X from mom, one X from dad. (Examples- hemophilia, colorblindness, deafness)
genetic drift
A change in the allele frequency of a population as a result of chance events rather than natural selection. (no selection, random effects through sampling variation can produce changes in allele frequencies, particularly prevalent in small populations, can cause small isolated populations to become genetically distinct from larger ones)
bottleneck effect (genetic drift)
when a large population is reduced in size and alleles are lost, variation is reduced. (cheetas as an example)
continuous characters
characters that have continuous distributions
continuous variation
range of small differences in a shared trait
(bell curve for distribution)
variation is difficult to eliminate completely through:
natural selection, rare deleterious alleles tend to be maintained at low levels
correlated characters
when individuals with a particular variant of one character also tend to have a particular variant of another character
if there is a strong selection on one trait, other traits may het "dragged" along with it
Other constraints of evolution
Evolution is also constrained by the laws of physics and chemistry
Surface area to volume ratios
DNA
deoxyribonucleic acid, a molecule in the cell containing genetic information. it is the "backbone" of sugar and phosphate molecules. attached to the sugar is of of 4 "bases."
scientific theory
a well-tested explanation for a set of observations or experimental results
Darwin vs. Lamarck
Lamark - Individuals lose characteristics that they do not use, and develop further on characteristics that they use a lot and that these changes would be inherited
Darwin - Some genetically programmed traits increase an organism's chances of surviving, do survive better, and spread through the population
process of evolution by natural selection
variation in heritable traits within a population
+ differential survival and/ or reproduction
= change in traits in a population over time
Continuous vs. Discontinuous variation
Continuous: Individuals vary within a range, generally due to multifactorial inheritance (e.g height)
Discontinuous: Two or more distinct categories of which a person can only fall into one, generally due to polygenetic or monohybrid inheritance (e.g blood type)