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Behavioral Ecology
The study of how organisms interact with their environment through behavior.
Ethology
The study of animal behavior mechanisms and the evolutionary significance of behavior.
Behavior
The observable actions of an organism in response to its environment, usually carried out by the muscular or hormonal system.
Proximate Causes
The stimuli and mechanisms triggering the behavior.
Ultimate Causes
The evolutionary explanations for why the behavior exists.
Nature vs. Nurture
Nature is the influence of inherited genetic programming.
Nurture is the influence of environmental factors.
Innate Behaviors
Behavior that is genetically controlled and does not require learning. It is often crucial for survival and found in nearly all organisms of a species.
Fixed Action Patterns (FAPs)
an innate behavior
A sequence of unchangeable, instinctive behaviors that is triggered by a sign stimulus, an external cue.
Once started, it is usually carried to completion.
Ex. A goose retrieving an egg back to its nest.
Behavioral Rhythms
Circadian, Circannual, Lunar
Circadian Rhythm: a daily cycle of rest and activity
Circannual Rhythm (changing seasons): mainly affects migration and reproduction, with common seasonal cues like daylight and darkness
Lunar Cycles: affect tidal movements
Migration
an innate behavior
The seasonal movement of organisms, usually triggered by environmental cues like temperature and daylight.
Signals and Communication
an innate behavior
Nocturnal vs Diurnal
Signals are behaviors that cause a change in another animal’s behavior.
Communication is the transmission and reception of signals.
Organisms use different signals to communicate:
Visual (warning coloration in frogs)
Auditory (bird songs, whale calls)
Tactile (grooming in primates)
Electrical (electric fish communication)
Chemical (pheromones used by ants and bees)
Nocturnal animals depend on olfactory and auditory communication while Diurnal animals use visual and auditory communication.
Kinesis
an innate behavior
A non-direction response to a stimulus.
Ex. Increased activity in pill bugs in humid environments
Taxis
an innate behavior
Phototaxis, Chemotaxis, Geotaxis
A directed movement toward or away from a stimulus. Positive taxis is toward the stimulus and negative taxis is away from the stimulus.
Phototaxis: in response to light
Chemotaxis: in response to chemicals
Geotaxis: in response to gravity
Learned Behaviors
Behaviors that develop through experience and environmental interaction.
Imprinting
a learned behavior
Learning that occurs during a critical period and is often irreversible.
Spatial Learning
a learned behavior
Recognizing landmarks to navigate the environment.
Cognitive Maps
a learned behavior
A mental representation of the surroundings for navigation.
Associative Learning
a learned behavior
Classical Conditioning and Operant Conditioning
Associating one stimulus with another.
Classical Conditioning: an arbitrary stimulus is associated with a reward or punishment
Operant Conditioning (trial-and-error learning): an animal associating one of its behaviors with a reward or punishment
Cognition and Problem Solving
Cognition: a process of knowing that may include awareness, reasoning, recollection and judgement
Problem Solving: the process of devising a strategy to overcome an obstacle
Social Learning
a learned behavior
Culture
Learning through observing and imitating others.
Culture is formed by social learning. It is a system of information transfer through observation and teaching that influences the behavior and fitness of individuals in a population.
Foraging Behavior
according to the Optimal Foraging Model
Efficient food-seeking strategies that maximize energy intake while minimizing risks and energy loss.
Sexual Dimorphism
Differences in the appearances of male and female organisms in the same species resulting from sexual selection.
Intersexual Selection
a type of sexual selection
Female Choice and Mate-Choice Copying
Members of one sex choose mates based on certain traits.
Female Choice: females drive sexual selection by choosing mates with specific behaviors and anatomical features that often correlate with health and vitality
Mate-Choice Copying: a behavior where individuals copy the mate choice of others
Intrasexual Selection
a type of sexual selection
Male Competition
Competition between members of the same sex for mates.
Male Competition: Reduces variation in males. This may involve agonistic behavior, often a ritualized contest that determines which competitor gains access to a resource.
Mating Behavior
Seeking or attracting mates, choosing mates, competing for mates, and caring for offspring.
Monogamy Mating Systems
A mating system where one sex takes one partner of the other sex.
Usually low sexual dimorphism (male and females look the same)
Polygamy Mating Systems
Polygyny and Polyandry
A mating system where one sex takes multiple partners of the other sex.
Usually high sexual dimorphism.
Polygyny: one male mates with many females, and the males are usually more showy and larger than the females
Polyandry: one female mates with many males, and the females are usually more showy than the males
Paternal Certainty
Internal and External Fertilization
Paternal certainty affects parental care and mating behavior.
Internal Fertilization: Paternal certainty is low because mating and birth are separated over time. Parental care is more likely to be by females.
External Fertilization: Paternal certainty is high because egg laying and mating occur together. Parental care is equally likely to be by males as by females.
Game Theory
Frequency-Dependent Selection
Evaluates alternative strategies where the outcome depends on each individual’s strategy and the strategy of other individuals.
Like rock-paper-scissors, each strategy will outcompete one strategy but be outcompeted by the other strategy. The success of each strategy depends on the frequency of all of the strategies, driving frequency-dependent selection.
Inclusive Fitness
The total effect an individual has on proliferating its genes by producing offspring and helping close relatives produce offspring.
Altruism
a cooperative behavior
Cooperative behavior that benefits other organisms at a personal cost.
Reciprocal Altruism
an altruistic behavior
Altruistism towards unrelated individuals that can be adaptive if the aided individual returns the favor in the future. This is limited to species with stable social groups where individuals meet repeatedly and cheaters who don’t reciprocate are “punished”.
Kin Selection
an altruistic behavior
Altruistic behavior that benefits relatives, increasing shared genetic fitness.
Hamilton’s Rule
A quantitative measure that predicts when natural selection will favor altruistism among related individuals, accounting for B, benefit to the recipient, C, the cost to the altruistic, and R, the coefficient of relatedness or fraction of shared genes.
Plant Behavior (Phototropism, Photoperiodism)
Physical and Chemical Mechanisms
Phototropism: Growth in response to light stimulus
Photoperiodism: Changes in physiological activity based on seasonal light availability
Physical Defenses: Thorns, bark and trichomes help protect against herbivores
Chemical Defenses: Toxins and distasteful compounds deter predators
Ecosystem
All the biotic(living) and abiotic(non-living) factors in a given area.
Metabolic Rate
The amount of energy an organism uses over a period of time.
Inversely proportional to body mass(smaller animals have higher metabolic rates)
Endotherms
Warm-blooded animals that maintain a constant internal temperature and require more energy.
Ectotherms
Cold-blooded animals that rely on environmental heat sources, using less energy.
The Energy Pyramid
A representation of how energy moves through an ecosystem via trophic levels.
Primary Producers: Autotrophs that create energy through photosynthesis(sunlight) or chemosynthesis(chemicals)
Primary Consumers: Herbivores, heterotrophs, that eat producers.
Secondary Consumers: Heterotrophs that eat herbivores.
Tertiary Consumers: Heterotrophs that eat other secondary or tertiary consumers.
Decomposers: Heterotrophs that break down dead organic material and recycle nutrients.
Food Webs
A diagram that shows the transfer of energy through an ecosystem using arrows(pointing in the direction of the energy flow).
Primary Production and Secondary Production
Gross Primary Production (GPP) is the total energy converted into chemical energy.
Net Primary Production (NPP) is the energy available to consumers after autotrophs use some for respiration.
Secondary Production is the energy transferred to higher trophic levels.
Only 10% of energy is passed between trophic levels, most is lost as heat.
Biogeochemical cycles
Cycles through which matter moves in ecosystems.
Water Cycle: includes evaporation, condensation, precipitation, groundwater flow
Carbon Cycle: moves carbon between the atmosphere, organisms, and fossil fuels, influencing climate change
Nitrogen Cycle: converts atmospheric nitrogen into usable forms through fixation, assimilation, and decomposition
Phosphorus Cycle: Vital for DNA and ATP, cycles through rocks, water, and living organisms
Population (and Population Ecology)
A group of individuals of the same species in the same geographical area.
Population Ecology is the study of the change in population due to births, deaths, immigration and emigration.
Density
The number of individuals per unit area.
Populations exhibit different patterns of Dispersion:
Clumped: Organisms gather in patches, often due to resource availability
Uniform: Organisms are evenly spaced apart, often due to territorial behavior
Random: No predictable patter
Life Tables
Survivorship Curves
Life Tables track survival patterns over time.
Type I: High survival in early/middle life
Type II: Steady mortality rate
Type III: High early mortality rate
Population Growth Models
Exponential (J-curve) and Logistic (S-curve)
An exponential curve shows rapid growth and occurs in ideal conditions.
A logistic curve shows growth slowing as it reaches the carry capacity and occurs when density exceeds resource availability.
Life History
The schedule and traits that affect an organism’s schedule of reproduction and survival.
This depends on
when reproduction begins
how often the organism can reproduce
the number of offspring produced per reproductive episode
K-selection (density-dependent selection)
a reproductive strategy
Produces few offspring, high parental investment
This is sensitive to population density and seen in high density populations that are close to carrying capacity (K).
R-selection (density-independent selection)
a reproductive strategy
Produces many offspring, low parental care
Density-Dependent Factors
Factors that affect the population more as population density increases
Competition for Resources: competition for limited resources increases (food, space, water)
Predation: denser populations attract more predators
Disease: Diseases spreads quicker because individuals are in closer proximity
Parasitism: Increase in parasites spreading among individuals
Density-Independent Factors
Factors that affect the population regardless of population density
Weather (storms, droughts, heat waves)
Natural Disasters (floods, fires, earthquakes)
Human Activities (Habitat destruction, pollution, deforestation)
Community
The group of different species that coexist and interact in a certain geographical area.
Habitat
The physical location where an organism lives.
Niche and Ecological Niche
All species in a community have a specific role, or niche: the full range of conditions and resources that an organism needs to survive, reproduce, and thrive.
The ecological niche is the role a species plays within its habitat, including its interactions with other species and its use of resources.
Fundamental Niche vs Realized Niche
Fundamental Niche: the potential range of conditions a species can tolerate and the resources it could theoretically use in the absence of competition and limiting factors
Realized Niche: the actual conditions and resources a species uses, considering ecological pressures such as competition, predation and disease
Competition
an interspecific interaction
when two species vie for the same resources, such as food, space and light
Competitive Exclusion Principle
Two species that occupy the same niche cannot exist indefinitely if resources are limited, as one will outcompete the other.
Niche Partitioning
The division of resources to minimize direct competition.
Through this, species can coexist by utilizing different resources or utilizing them at different times or in different ways.
Predation
an interspecific interaction
Cryptic Coloration, Batesian Mimicry, Mullerian Mimicry
The interaction where one species, the predator, hunts and consumes the other, the prey.
This relationship drives the evolution of defensive traits in prey species:
Cryptic Coloration: prey species blend into their surroundings to avoid detection by predators
Batesian Mimicry: a harmless species resembles a dangerous or poisonous species to deter predators
Müllerian Mimicry: two harmful or poisonous species resemble each other to reinforce predator avoidance
Herbivory
a type of predation
The interaction where herbivores feed on plants.
This relationship drives evolution of traits in both plants and herbivores.
Plants evolve physical defenses like thorns or produce toxins. Herbivores may evolve mechanisms to counter these defenses.
Symbiotic Relationships
Facilitation, Parasitism, Mutualism, Commensalism
Interactions where two or more species live together in close association
Facilitation: at least one species benefits and neither are harmed
Parasitism: one species benefits at the expense of another, typically harming the host species
Mutualism: both species benefit
Commensalism: one species benefits and the other is unaffected
Species Diversity
The variety and abundance of different species within a given ecosystem or community.
Simpson’s Diversity Index (D)
A mathematical formula used to measure the species diversity of a community, taking into account the number of species and the relative abundance of each one.
The higher the value (closer to 1, further from 0), the greater the diversity.
Invasive Species
Non-native species that are introduced to an ecosystem and cause harm by outcompeting local species, altering habitats and disrupting ecological processes.
They reduce species diversity and ecosystem stability.
Keystone Species
A species whose role in an ecosystem has a disproportionately large effect on the structure and diversity of that ecosystem.
They maintain species diversity by preventing any species from becoming overly dominant.
Ecological Disturbances
Events or processes that disturb the structure of an ecosystem.
Ex. Natural disasters, Human Activities
Ecological Succession
The process by which ecosystems develop over time following an ecological disturbance, involving the gradual replacement of species in a community resulting in a more stable, diverse ecosystem.
Primary Succession
an ecological succession
Occurs in areas where ecosystems did not previously exist, such as after a glacial retreat or volcanic eruption.
Begins with pioneer species like lichens and mosses.
Secondary Succession
a type of ecological succession
Occurs in areas where an existing ecosystem was disrupted but soil and some organisms remain, such as after a forest fire or abandoned agricultural land.
Due to the presence of soil and nutrients, this is faster than primary succession.
Human Activities
a major cause of…
A major cause of biodiversity loss and ecosystem disturbance.
Habitat Loss: the destruction of natural habitats due to human activities, causing species to lose their homes and die off
Overharvesting: the excessive harvesting of natural resources depleting species populations
Global Change: climate change and weather patterns can affect temperature, rainfall and resource availability for species
Biogeographic Factors
on the distribution of species across the globe
Latitude: Biodiversity increases closer to the equator because tropical regions typically have more diversity than temperate or polar regions
Area: Larger areas support more species because they provide more habitats and resources. Island ecosystems have fewer species than continental areas.
Pathogens: The spread of diseases drastically affects species populations, especially when species lack immunity or are in stressed environments.