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Charles Darwin
-formulated theory of evolution/natural selection
Darwin's three postulates
1. the ability for a population to expand is infinite, but the ability for an environment to support populations is always finite
2. organisms within a population vary, and this variation affects the ability of individuals to survive and reproduce
3. variation is transmitted from parents to offspring
morphology
organisms size, shape, and composition- can change over time so that the organism can adapt to their environment
Gregor Mendel
-"father of modern genetics"
-experimented with pea plants (color-green or yellow, & texture- wrinkled or smooth) which eventually led scientists to discovery of alleles
alleles
different varieties of a single gene (can be dominant or recessive)
homozygous
having two identical alleles for a particular gene
heterozygous
having two different alleles for a particular gene
genotype
an organism's genetic makeup, or allele combinations
phenotype
physical characteristics of an organism
4 characteristics of natural selection*
1. sexual reproduction produces VARIATION
2. some offspring are better suited to environmental conditions than others
3. those that are better suited to environmental conditions will produce more offspring than others (fitness= reproductive fitness)
4. over many generations there is enough change produced that a new species is produced
all species are a combination of:
1) primitive traits
2)derived traits
primitive traits
adaptations of the past (ex. pentadactyl)
derived traits
adaptations to recent timed (ex. bipedalism)
what are the 5 primate characteristics?
Primitive body plan
Dental formula
Grasping hand and foot
Increased reliance on sight
Less reliance on smell
K selected
what does primitive body plan consist of?
-one bone in upper limb
- two bones in lower limb
-five digits on hands and feet
what is a primate's dental formula?
2-1-2-3 or 2-1-3-3 (incisors-canines-premolars-molars)
hederodant
hederodant
dental formula of primates, teeth are in different positions in jaw and have different functions
homodant
dental formula found in reptiles, teeth all have same shape and function
2-1-2-3 dental formula
old world monkeys, apes humans
2-1-3-3 dental formula
prosimons, new world monkeys
K versus R selected
- K selected: few offspring, large investment
- R selected: many offspring, small investment
altricial young
offspring that is completely helpless after birth and dependent on their parents
precocial young
offspring born in a relatively mature state, eyes open
where do most primates live?
in the tropics, near equator
arboreal
living in trees (nearly all primates are arboreal)
biological species concept
if two animals can interbreed, they're part of the same species
arboreal hypothesis
the proposition that primates' unique suite of traits is an adaptation to living in trees:
1. grasping hands and feet
2: stereoscopic vision
ape characteristics
larger body size, longer lifespan, longer interbirth interval, upright posture, larger brain, shorter muzzle, no tail, increased reliance on vision
function of incisors
cutting food/taking a bite
function of canines
tearing food/killing your meal/defense and display (bigger in males)
function of premolars and molars
grinding and crushing
bunodont molars
low crowned, rounded cusped molars present in frugivores or omnivores
hypsodant teeth
(seen in horses) high crowned, used especially for grass or leaf eating
lophodant or selenodant
complex cusps, used especially for grass or leaf eating
carnassials
slicing molars and premolars present in carnivores
frugivore
fruit eaters
-big incisors
-bunodont molars
-short gut
folivores
leaf eaters
-small incisors
-sharp, high/tall crowned molars &premolars
-long gut
carnivore
meat eater
-small incosors
-large canines
-carnassials
-short gut (meat= high quality, dense in protein)
omnivore
(generalists - eat anything)
-large incisors
-bunodont molars
-canines variable
-short to medium gut (humans known for short gut)
Milutin Milankovitch
scientist who discovered astronomical forcing of global climate
steps of astronomical forcing of global climate
1.changes in shape and orientation of earth's orbit
2. differences in distance and angle between earth & sun
3. changes in amount of solar radiation reaching earth
4. changes in global climate
Linnaeus
proposed a new system of organization for plants, animals, and minerals, based upon their similarities- binomal nomenclature
-taxonomy (classification of organisms)
binomal nomenclature
the naming system for organism in wich each organsim is given a two part name-a genus name of different species in an area
(ex. homo sapiens
uniformitarianism
a theory based on the work of James Hutton and made popular by Charles Lyell in the 19th century. This theory states that the forces and processes observable at earth's surface are the same that have shaped earth's landscape throughout natural history.
microevolution
small changes within a population over a short period of time (change in gene frequemcies
macroevolution
large changes within a population over a long period of time (extinction or creation of new species)
7 mechanisms of microevolution that change allele frequencies
1)natural selection
2)mutation
3)genetic drift
4)population bottleneck/ Founder Effect
5)assortative mating
6)inbreeding
7)gene flow (migration)
what is mutation?
change in DNA sequence, appearence of new alleles
what is genetic drift?
variation in the relative frequency of different genotypes in a small population, owing to the chance disappearance of particular genes as individuals die or do not reproduce.
what is the founder effect?
shows that allele frequencies in small founding population may not represent those in parent population
what is assortative mating?
when individuals mate with those who are similar to themselves
what are the effects of inbreeding?
more likely to look alike & inherit double copies
what is gene flow/migration's microevolutionary effect?
population movement introduces new alleles
4 kinds of macroevolution evolutionary events
1) stasis
2) extinction
3) anagenesis
4) cladogenesis
stasis in macroevolution
species lineage shows NO change
extinction
termination of species
anagenesis
gradual modification of form over evolutionary time, without branching speciation
cladogenesis
BRANCHING SPECIATION the formation of a new group of organisms or higher taxon by evolutionary divergence from an ancestral form.
allopatric model of speciation
(other land model) species becomes isolated geographically and changes in its own way as it evolves
sympatic model of speciation
species live in same area but use different resources (niche partitioning) ex. finches
isolating mechanisms include
1) morphological differences
2) genetic incompatibility
3) behavior
4) mate recognition (SSMRS)
examples of isolating mechanism behavior
- posture: exhibits willingness to mate
-vocalizations
convergent evolution
process whereby organisms not closely related (not monophyletic), independently evolve similar traits as a result of having to adapt to similar environments or ecological niches.
convergent evolution example
the wings of bats and birds
DNA hybridization
the process used to compare structural characteristics of organisms to determine how closely related they are
DNA hybridization process
1. extract DNA
2. cut double helix strands by heating
3. reform double helix by connecting the different DNA into "hybrid strands"
4. determine melting temp and calculate difference
molecular clock
model that uses DNA comparisons to estimate the length of time that two species have been evolving independently
fossil
mineralized remains of animal or plant (usually bones or teeth)
trace fossil
sign of animal's or plant's interaction with the environment (ex. footprints or burrows)
artifact
any object made or modified by hominins (ex. stone tools)
types of rocks
1. volcanic rocks
2.sedimentary rocks
3. metamorphic rocks
volcanic rocks
derived from molten crust
-dateable by potassium/argon and argon/argon dating
-source for toolstone
sedimentary rocks
composed of particles of other rocks (ex. sandstone)
-contains fossils or artifacts
metamorphic rocks
sedimentary rocks transformed by heat or pressure (ex. marble)
kinds of volcanic rock
1. magma - underground
2. lava - extruded at surface
3. ash - exploded into air (tephra or tuff, pyroclastic flow)
volcanic ash (tuff, tephra)
records single instant in time
-dateable by K/Ar & Ar/Ar
- excellent for correlating among sites
loess
(ex. of sedimentary rock) Eolian deposit- windblown silt
particle size in sedimentary rocks
reflects energy in sedimentary environment (how much energy is required to transport particles)
mudstone,shale -> sandstone -> conglomerate (pebbles, cobbles, boulders) ..... low energy -> high energy
law of superposition
Nick Steno- law that states the layers at the top are the youngest and those on the bottom are the oldest layers (strata)
lithostratigraphic vs biostratigraphic correlation
lithostratigraphic: strata collected based on rock type
biostratigraphic: strata collected on basis of fossils they contain
radiocarbon dating
-type of radioactive dating measures decay of 14C into 14N
-half life: 5730 yrs
-approximate limit of 50,000 years
argon-argon dating
A high-precision method for estimating the relative quantities of argon-39 and argon-40 gas; used to date volcanic ashes that are between 500,000 and several million years old
when did africa and eurasia connect?
17 ma
when did continents reach modern position?
17 ma
common isotopes of oxygen
18O - in cold periods, sea water is higher in 18O (heavy)
16O- ice takes up 16O (lighter)
Milankovitch effects
changes in:
-temperature
-rainfall
-vegetation
-glaciers
-sea level
what is gene flow/migration's microevolutionary effect?