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directional selection
changes avg phenotype in pop in one direction
reduced genetic diversity since 1 phenotype is favored
purifying selection
shifting to one extreme

disruptive selection
intermediate phenotypes are selected against and extreme phenotypes are favored
increases amount of variation
has the opposite effect on stabilizing selection

stabilizing selection
reduces both extremes and favors the average
reduces the amount of variation
gets skinnier in the middle

balancing seelction
occurs when no singles allele has a distinct advantage
heterozygote advantage and frq dependent selection

heterozygote advantage
individuals have higher fitness than homozygous individuals
sickle cell
frq dependent selection
certain alleles are favored when rare
not common
life history describes
growth, development, reproduction, and survival
life states defined by
embryo, juvenile, and adult stages across tree of life
what environmental factors influence state length and reproductive output
fecundity
parity
parental investment
longevity/life expectancy
fecundity
the number of offspring per reproductive episode
parity
number of episodes of reproduction
parental investment
energy incurred by parent to raise offspring
longevity/life expectancy
lifespan of the organism
why do we see variation in life history
growth, maitenance, reproduction

growth
body size
maitenance
lifespan
reproduction
number of offspring within a reproductive bout and the frq of reproduction
principle of allocation
only a limited amount of energy that is distributed to the 3 pts (reproduction, maitenance, growth)
fast-slow continuum
reprsents 2 extremes of how tradeoffs manifest
often reflect variation in life form, habitat, or environmental condition
fast side
greater number of offspirng, shorter life span, faster growth, earlier reproduction, earlier sexual maturation, smaller parental investment
slow side
fewer offspring, longer lifespan, slower growth, delayed reproduction, later sexual maturation, greater parental investment
life history and pop growth

semeiparity
organisms that reproduce once
mayflies
interoparity
organisms that reproduce multiple times in their life
iterative
oak trees
costs and benefits of mice reproduction (6 pups every 3 wks)
costs: low prenatal care, increased number of offspring, decreased life span
benefits: increased gene pool with mating patterns, at least one will survive
cost and benefit of tortice reproduction (2-16 eggs per year)
costs: takes longer to reach sexual maturation, decreased number of offspring
benefits: increase prenatal care, longer lifespan
Offspring v. prenatal care: 1947 study
modified eggs in European magpie nest
manipulated total offspring without sacrificing parental services
7 eggs is the perfect amount bc most wil survive and it is the perfect amount to provide for
determinate growth
growth pattern in which an individual does not grow anymore once it initiates reproduction
inter-determinate growth
growth patterns in which an individual continues to growth after initiating reproduction
population
group of species that live in the same area and inter breed
darwins quote highlights how
there are many factors in place that pops are diverse and numbers are kept in check
Factors contributing or detracting from pop growth
births (+)
deaths (-)
immigration (+)
emigration (-)
exponential growth
consistent high amounts of growth
represents continuous growth, but not all organisms have a continously growing pop

exponential growth equation
r=intrinsic rate of rate
per-capita growth rate (births-deaths) under ideal conditions

rate of pop growth for exponential growth
r=net number of offspring individual time

geometric growth model
dots represent organisms that ahve 1 distinct breeding event per yr
compares pop numbers at set intervals
doesnt assume continuous births and deaths in a pop

geometric growth equation
lambda is the ratio for a population in 1 yr to the preceding yr

Populations with discrete reproductive periods increase by
geometric from
equation for reproductive periods from one time to the next

equation for repoductive periods over long time periods

how is lambda different from r in exponential growth
when discussing discrete growth patterns, lambda compares year with previous yrs
at different time pts we can solve for lambda
when more deaths than births, see
lambda decrease

when births and deaths are the same
flat line

when more births than deaths,
increasing lambda

lambda can also be calculated for
exponential growth we just need to know N at 2 pts in time
what factors limit pop growth
density dependent factors
negative density dependence
when pop growth decreases and pop density increases (from increased competiotion)

positive density dependence
when more indvidual present result in increased fitness of the pop, resulting in increased growth
often observed in really small pops

density independent facotrs
factors other than pop density determine pop size (abiotic factors)
temp and precipitation
negative density dependence
an ecological process where a population's growth rate or individual fitness increases as population density increases
Negative influences on pop
competition ,limited resources
positive influences on pop
increase surplus, increased interaction, start low pop #
Consequences of low pop density
difficulty in finding mates
poorer predator detection
lower genertic diversity, inbreeding
populations can experience
positive or negative density dependence at different times
logistic (sogmodial) growth model
what is the equation
assumes all individuals are the same

carrying capacity
amount of resources that can support a pop
represented by a pop number
can change over time
reindeer on st paul island
original carrying capcity reduced by overgrazing of lichens
reindeer switched to grass and grass roots
current carrying capacity: around 400 reindeer
facotrs for pop growth and density for barnicles
food, salinity, water, density/space
age structure is important to pop growth
populations with high proportions of younger individuals (of reproductive age) can have higher growth rates
metapopulation
composed of subpopulations that are connected to eachother
metapopulation models
black dots are occupied; open dots are not
Lines indicate migration; dashed lines outline indicate high migration
Levins model
patches are either occupied or unoccupied; otherwise equal quality of patches to support population growth

Mainland model
mainland patch is highly occupied because it is a higher quality area.
Contribute to other, low quality patches

patchy population model
different quality of patches, but see migration between; constant rates of high migration among all patches

Non-equilibrium model
migration between some patches, but not all

rescue effect
the phenominon of dispersers supplementing a declining subpopulation that is headed toward extinction
effects of patch size and patch isolation
The shrew was much less likely to occupy patches that were small or more isolated.
The skipper butterfly in Britain occupies the largest patches of grasslands and those that are the least isolated.
The line indicates the combinations of patch area and patch isolation that correspond to 50 percent probability of patch occupancy.

Consumer resource interactions
interactions that account for organisms being a resource for another organism OR consuming another organism
Competition
A relationship where organisms fight for the same limited resources, such as food, water, space, or mates.
Predation
A relationship where one organism (the predator) hunts, kills, and eats another organism (the prey).
parasitism
A relationship where one organism (the parasite) benefits while the other organism (the host) is harmed, usually without being immediately killed.
herbivory
A relationship where an animal eats plants or algae.
herbivores
consume plant and plant materials
mesopredators
consume herbivores, often smaller
top predators
consume herbivores and mesopredators
parasitoids
specialized predators
live inside and consume their host during development
parasitoid wasps
large group of wasps that need to lay their eggs inside other insects to complete their lifestyle
useful in contorlling both niative pests and invasive species
do not sting or bite people
California redscale
atiny insect with a hard shell
feeds on the sap of citrus trees and other plants.
The parasitoid wasp lays its eggs under the hard shell of the scale insect. When the eggs hatch, the larvae slowly consume the scale insect.
After introducing large numbers of scale insects to trees the scale insect population grew rapidly. Soon after the addition of the scale insects, the number of juvenile and adult parasitoid wasps began to rapidly increase. This increase in parasitoid wasps caused a subsequent rapid decline in scale insects.
lotka-volterra model
a model of predator-pret interaction that incorporates oscillations in the abundances of predator and prey pops and shows predator number logging behind these of their prey
prey pop change lotka-volterra model
rn= growth rate x number of prey
cnp= capture efficieny, np-random encounter with predator
