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Scientific theory
A broad, well tested explanation with predictive value (leads to many accurate predictions)- like gravity
Evolution
The change in organisms throughout earths history; change in a populations genetic composition over generations
Is evolution just a theory
No it’s a scientific theory, backed by a lot of evidence
Do individuals evolve
No populations do
Can an organism evolve within its lifetime
No
Does evolution explain the origin of life
No; only describes how existing populations of living things change and diversify over time. For biological evolution to occur, life must already exist
Did organisms evolve on purpose
No, would suggest a goal oriented process; cannot influence the evolution of its own structures in response to the environment
Are species always evolving into higher or better beings
No, they just adapt to changes in the environment
Does evolution create new life forms by dramatic mutations
No, it occurs generation to generation and is passed to offspring; DNA cannot produce new structures in its lifetime
Is evolution a completely random process
No; The presence of certain genes/traits allows the population to adapt and survive and reproduce
Aristotle and evolution
Viewed species as unchanging/fixed based on an increasing ladder of complexity (scale naturae)
Age of reason
Pre Darwin; the rise of scientific inquiry and the use of reason; emphasis on observation, experimentation, and challenging established doctrines
Linnaeus
Created an orderly classification system, grouping species into increasingly general categories, father of taxonomy, invented binomial naming
Binomial naming
Made by Linnaeus, 2 parts: genus (homo) and species (sapien) all italicized
James Hutton
Geologist, gradualism theory: earths geological features (canyon, valley, etc) are a result of gradual mechanisms, slow continuous processes
Charles Lyell
Father of modern geology; expanded huttons ideas by introducing uniformitarianism ; wrote principles of geology; earth is extremely old
Uniformitarianism
By Charles Lyell; same geologic processes in past as today, same rate as today
Lamarck
Linked evolution to adaptation via changes in environment; extinct species have been replaced by descendants with new features; first to propose a mechanism for evolution: use it or lose it; also believed in inheritance of acquired characteristics (wrong)- if you lose an arm your child will lose an arm
2 main ideas of Darwin’s theory
descent with modification and natural selection
descent with modification
Darwins theory; species that live today are descended from ancestral species that were different from the present day species
natural selection
Darwins theory; survival of the fittest; reproduction of individuals with favorable genetic traits that survive environmental change- leads to evolutionary change; heritable variation exists in most species, more offspring are produced than can survive so there is competition for resources
does evolution lead to perfection
no, mutations are random and it often takes many to lead to a new trait which can help or harm or neither; evolution is change in organisms over time
direct observation
evidence of evolution; observe then infer; soapberry bugs get beak length that matches depth of balloon vine fruit
fossil record
evidence of evolution; whales are mammals and their ancestors lived on land- shows transition of adaptive form over time
homologous structures: divergence
evidence of evolution; forms related by common ancestry (like human, dog, bird, whale arms); similarity in layout and construction but the functions are different; indicate that a species is diverging from its ancestors
developmental homology
type of homologous structure; fish and human embryo are very similar, meaning we share a common ancestor
vestigial structures
evidence for evolution; remnants of ancestral (homologous) structures with no present adaptive function (wisdom teeth, tailbone, etc); blind cave salamanders have eyes but dont use them bc they descended from species that could see
analogous structures
structures with similar functions but different ancestors; sometimes misleads us when trying to trace back to common ancestor
convergence
evidence for evolution; unrelated species have similar adaptations under similar environmental conditions ; torpedo shape for swimming- live in same environment so develop similar adaptations; results in analogous structures; wings in birds vs insects
biogeography
distribution of species which corresponds to geographic history; South America is nearest mainland to Galapagos- marine iguana in Galapagos ocean and tree dwelling in South America; they are most similar bc marine evolved from tree dwelling
molecular biology
evidence of evolution; closely related organisms have similar DNA; genetic code is universal, so all of us come from a common ancestor
species
basic unit of biological classification; group of organisms that can interbreed and produce viable, fertile offspring
morphological species
same species based on what they look like; white oak in Tennessee vs texas look alike
biological species
set of populations whose members potential interbreed in nature to produce fertile offspring and do not successfully interbreed with other groups; share the same gene pool and there is gene flow
ecological species
set of organisms adapted to a specific set of resources; morphologically similar, diff species based on utilization of food source; bacteria
gene pool
sum of all alleles in the population
gene flow
movement of genetic material
what if there is no gene flow between two populations
they are different species
how are different species reproductively isolated from other species (biological species concept)
via natural biological barriers: prezygotic and posyzygotic
pre zygotic biological barriers
prevents fertilization (meshing egg and sperm together); 5 types: temporal, habitat, behavioral, gametic, mechanical
post zygotic biological barriers
reproduction happens and causes egg and sperm hybrid but there is a barrier formed after; 3 types: reduced hybrid viability, reduced hybrid sterility, hybrid breakdown
habitat (ecological) isolation
pre zygotic barrier; never meet bc they live in diff habitats
temporal isolation
pre zygotic barrier; can live in same area but breed at diff times of the year
behavioral isolation
pre zygotic barrier; have diff mating rituals/calls/dances; presence or absence of certain behavior prevents mating (like bird sounds)
mechanical isolation
pre zygotic barrier; incompatibility of the sexual organs
gametic isolation
pre zygotic barrier; when gametes cannot fuse together due to difference in gamete cells (wrong pollen on a flower)
gametic isolation
pre zygotic barrier; sperm of one species may not be able to fertilize eggs of another species; proteins on sperm will only bind to specific receptors on the covering of the egg in order to fertilize
reduced hybrid viability
post zygotic barriers; embryo fails to develop or is weak and dies
reduced hybrid fertility
post zygotic barriers; hybrid survives but is sterile or almost so; horse (64) and donkey (62) mate but chromosomes dont pair up properly during meiosis so offspring becomes infertile
hybrid breakdown
post zygotic barriers; 1st generation offspring are viable and fertile but when they mate with each other offspring is feeble or sterile; so 1st get hybrids are good but 2nd gen isnt
biological species cannot apply to fossil species why
reproductive information doesn’t fossilize
biological species cannot apply to asexual species why
no use of zygotes or hybrids
microevolution
how new species form; changes over time in allele frequencies in a population; any change in population allelic or genotypic frequency over time; evolution on smallest scale
how do new species form
micro and macro evolution
macroevolution
the processes that gave rise to new species and higher taxonomic groups with widely divergent characters
speciation
formation of new species; may occur with or without geographic separation; 2 types: allopatric and sympatric
population
group of individuals of the same species that live in same area and interbreed and produce fertile offspring
allopatric speciation
geographic isolation (w barrier); can happen 3 ways: dispersal, vicariance, adaptive radiation
dispersal
allopatric speciation; few members of species become isolated geographically for a long period of time and changes in each group over time leads to divergence; adapts to new environment and evolution occurs via natural selection
vicariance
allopatric speciation; natural situation arises to physically divide the organisms (mountain, river, etc)
hybrid zones in allopatric speciation
area where 2 closely related species interact and interbreed; 3 types: reinforcement, fusion, stability
reinforcement
hybrid zone; due to natural selection, hybrids are less fit than the purebreds, reinforcing the pre zygotic barriers; species continue to diverge until hybridization can no longer occur; A on top of V
fusion
hybrid zone; reproductive barriers weaken until the 2 species merge to become 1; diamond
stability
hybrid zone; barriers remain the same but fit hybrids continue to be produced; rectangle
Dodds experiment
allopatric speciation; flies raised on starch vs maltose; mated them; showed that there was a diet-developed mating preference
adaptive radiation
allopatric speciation; many adaptations evolve from a single point of origin, causing the species to radiate into new ones; from one founder species of bird, multiple others evolved, each with its own distinctive characteristics (beak shape changes relative to what it eats)
sympatric speciation
creation of new species; occurs in same geographical area; based on mating behavior (fish under different light causes interbreeding) and habitat differentiation (2 populations isolated by use of different resources, lay their eggs on certain fruit and mate there); 2 types of chromosomal errors during cell division: aneuploidy and polyploidy
aneuploidy
sympatric speciation; gametes have too many or too few chromosomes due to nondisjunction (chromosomes fail to separate) during meiosis; will have 2n+1 or 2n-1 chromosomes
polyploidy
sympatric speciation; cell or organisms has extra set or sets of chromosomes; results from error in meiosis; 2 types: autopolyploidy and allopolyploidy
autopolyploid
type of polyploidy; sympatric speciation; individual that has more than 2 chromosome sets that are all derived from a single species; chromosomes divide by cell doesn't , so go from 2n to 4n cell
alloploidy
type of polyploidy; sympatric speciation; 2 species mate to produce viable offspring; takes 2 matings to major viable fertile offspring
punctuated equilibrium
rates of speciation; species exhibit a large change in a relatively short period of time followed by long periods of freezing/stasis; remains unchanged for a long time
gradual speciation
species diverge gradually through time with small steps; multiple diff stages in time
without _____ evolution by natural selection cannot happen
genetic variation
phenotypic variation
mostly genetic; the difference in observable traits—such as size, color, or behavior—among individuals in a species; environment can influence expression, creating non-heritable variation; larvae fed on oak flowers and looks like them while larvae fed on oak leaves looks like twigs
genes and inheritance of genes leads to different types of
variation; 2 types: discrete genetic and continuous
discrete genetic variation
2 or more alleles at a single gene locus; a trait is one thing or another, no mixing
continuous variation
phenotypes produced by combined effects of 2 or more genes; controlled by multiple genes and environmental factors
sexual recombination
produces genetic diversity among offspring; crossovers, independent assortment, random fertilization
new alleles arise from
mutations in DNA; only mutations in germ cells that make gametes are passed down; point mutation or chromosomal alterations
does most DNA variability affect phenotype
no; no new allele bc protein translation/gene expression is not affected; non coding sequence is removed via splicing
harmful alleles
most are like this but they may be “hidden” in heterozygotes (Pp) but will pass it on; if environment changes they may become adaptive
neutral alleles
doesn’t provide any advantage or disadvantage; new phenotype doesn’t affect likelihood of leaving offspring; if environment changes they may become adaptive
population genetics
study of what changes allele frequencies in populations through time
genotypic frequency
% of each genotype (AA, Aa, aa) in the population
allelic frequency
% of each allele in the population (A, a)
hardy-weinberg equilibrium
describes a non-evolving population under ideal conditions (so evolution isnt necessary); if a large population reproduces randomly then the genetic frequencies dont change in the next generation (stay in eq); lets us detect microevolution- if the actual ratios dont equal the expected HW ratios then the population is evolving
hardy Weinberg conditions (all must be true)
no mutations, mating is random, no natural selection (equal survival), very large population size, no gene flow in or out
KNOW HOW TO DO P2+2PQ+Q2 EQUATION
p
frequency of dominant allele
q
frequency of recessive allele
p2+2pq+q2 = 1 ; p+q = 1
p2 = frequency of homozygous dom genotype CRCR ; 2pq = frequency of heterozygous genotype CRCW ; q2 = frequency of homozygous recessive genotype
mechanisms of microevolution
natural selection, genetic drift, gene flow
natural selection and microevolution
acts non-randomly on phenotypes- favors individuals with certain characteristics/traits; changes allelic and genotypic frequencies of populations non randomly; always leads to adaptation of population to current environment; flies carrying resistance to DDT
genetic drift and microevolution
genetic frequency changes due to random events; 2 types: founder and bottleneck effect; often occurs in small populations; can cause random changes in allele frequency, reduces genetic diversity, causes allele fixation
fixed allele
allele that is the only option that exists for that gene in a population (always homozygous for all members) all have brown coat BB and no white
founder effect
part of genetic drift; a few individuals become isolated from a larger population and start their own new population; less diverse founder population, causing more genetic drift and some adaptive alleles are lost; can cause maladaptive allele frequency increased (high rate of inherited blindness on an island)
bottleneck effect
part of genetic drift; an event drastically cuts population size due to sudden change in environment (flood, fire, etc); by chance, certain alleles are overrepresented by certain survivors and some are lost
gene flow and microevolution
alleles move in/out of the population; includes migration of adults, dispersal of gametes, seeds, larvae (western vs eastern deer population); adds genetic diversity, reduces differences between populations
relative fitness
contribution an individual makes to a gene pool of the next generation relative to the contribution of the other individuals; best reproductive success; brown Beatles leave more offspringg due to color so they have a higher fitness relative to the green onesdif