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Wetlands are highly productive because
high organic primary production and have alrge amounts of both animal and plant biomass
Wetlands regulate the climate through
carbon storage, methane production, and coal production
Nitrogen is abundant in ____ but scarce in ____
air, organisms
Nitrogen fixation
done by Cyanobacteria that converts atmospheric N2 to NH4
Denitrification
removal of N in plant tissues or organic material and is converted back to atmospheric N2
Nitrogen is limiting in _____ and phosphorus is limiting in ___
land, water
Sediment can store nutrient such as
N and P leading to peat build up
Wetlands can act as storage for
rare species, biologial diversity, and genetic diversity
Wetland services and management
individual species, large amounts of biodiversity, water purification, and huge economic value
Causes of flooding
rain, melting snow, natural rising and falling of water, natural or man-made disasters
Wetlands ____ flooding
reduce
Wetlands can record historical data including
climate data, organisms, succession, contaminates, and depostition rates
Flooding causes variations
within years and among years
Biological consequences of flooding
water fluctuations, breeding and maturing of offspring, behavior and life cycle of fish, macroinvertebrate composition, and endangering rare plants
Water level fluctuations depending on sources of water
Rivers → seasonal flooding
Lakes → rising and falling shorelines
Beaver ponds → dab building and collapse
Potholes/vernal pools → rain and snowmelt
Peatlands → more stable water tables
Flooding in swamps
zone woody plants at high elevation and occasional flooding allows vegetation to survive
Flooding in wet meadows
kills woody plants, plants can establish more rapidly, and requires occasional flooding
Flooding in Marshes
flooded for most of the growing season and plants are tolerant of flooding
Hydrological effects of dams
stabilized water levels, shifted food timing, increased flooding (levees concentrate flow), and decreased flooding (reduced floodplain area)
Upstream effects of flooding
water levels depend on dam purpose, extreme flucations are possiblem and reversed or extreme cycles can cut vegetation
Downstream effects of flooding
reduced spring floods (global loss of wetlands), conversion of wet meadows to woody vegetations), and loss of biodiversity and habitat

Predicting consequences of flooding
Stablizied water levels reduces plant diversity and marsh area
Six elements make up all organisms
CHOPNS
__ and __ are loved by plants and can be used to determine habitat suitability, also called ___
P and N, fertility
Fertility controls ___. Thus __ and __ supplies limit plant growth.
primary productivity, N, P
Infertile wetlands include ___ and are characterized by …
Pocosin (sandy peat soils and wood shrubs), relying on rain water( low nutrients), sensitivty to increase nutrient availability, and strong gradients in plant compostion
Infertile habitats have unique species including plants
that can tolerate nutrient shortages, have slow growth, evergreen foilage, nutrient storage, anti-herbivory defenses, and mycorrhizae
____ is the limiting factor is fertile habitats leading to plants that are tall and fast growing
light
Fertility varies in ____ in wetlands that can be used to explain the patterns of a wetland
gradients
Peatlands (fertility)
low in nutrients, pH and calcium concentrations play a major role, water supply, water level, and location play a role in species composition and growth
Mycorrhizae
fungi with a symbiotic relationship to plants roots, Mycorrhizae recieve sugars and plants recieve nutrients in return
Most plants products contain less than ___ N which is ___ for herbivores
5%, bad
Animal strategies for increasing Nitrogen
synchronzining life cycles with food availability, selecting tissues with higher N, eating quickly and digesting more efficiently, supplementing plant food with animal protein, and territoral and social behavior
Four general consquences of nutrient accumulation in wetlands
decreased biodiversity due to increased biomass causes by alleviation of nutrient limiations
increased nutrients cause plants to become more palatable to herbivore, increasing herbivory and herbivore numbers
increased floating algae can kill aquatice macrophytes
decomposition of algae and macrophtyes lead to increased O2 consumption and hypoxic conditions
Sources of nutrients
urband areas (point source), rural areas (diffuse source), and rainfall
Solutions to excess nutrients
Urban: wastewater treatment, reduce lawn fertilizer, clean up after pets
Rural: change agriculture and forestry practice and reduce road building
Precipitation: stricter environmental regulations of buring fossil fuels and reduce car usage
Calcium in Wetlands
reduces acidity, binds with P to form calcium phosphates lowering amount of P available to plants, and calcium concentrates and pH are very important for determing wetland type (Bogs: low pH and sphagnem moss)
Calciphilous
species restricted to calcium-rich soils
Disturbance
a short lived event that removes biomass and causes measurable change in the properties of an ecological community
Common distrubance examples
fire, ice scour, storms, herbivory
Distrubance can alter
biomass, substrate, and nutrients
4 properties of disturbance
duration (how long)
intensity (severity of the event)
frequency (higher intensity of disturbance =less frequent)
area (amount of land effected by distrubance)
Seed banks
buried reserves of viable seeds
Fire can
remove biomass inceasing light availability and ash contains P
Marshs and wet meadow species can only regenerate in ___ provided by ___
gaps, disturbance
Seeds can detect disturbace through
fluctuations in soil temp, increased light, and changes in the quality of light
Highest intensity disturbance is equals
highest species richness
Saline environments have ___ seed densities
lower
Colonization patterns from erosion and deposition
Herbaceos pioneer plants → small pioneer trees → mixed pioneer and successional species (closed canopy forms) → mature plant community
Low intensity fire disturbance
removes above ground biomass and shift species composition and wetland type
High intensity fire disturbance
burn organic soil, create new species composition, often create open water
The everglades are a ___ of different vegetation types due to ___ disturbance
mosaic, fire
Fire disturbance in peatlands can record history of disturbance and system response in ____ and ____ layers
charcoal, macrofossil
Feedback loops in peatlands with fire
fire → carbon release → higher temperatures → more frequent fires
Types of ice disturbance
ice foot formation, spring ice, riparian effects, and ice scour
Ice scour
large chunks of ice driven by wind and currents can rip out sediment and vegetation, removes fine sediment, lowers OM conten, and increases habitat without shrub coverage
Effects of wave action
moderate exposure: expand wet meadows and marshes
high exposure: sand or gravel shorelines
chronic exposure: removal of fine sediments
amplify ice effects: increase movement and erosion during freeze and thaw conditions
Animals ___ biomass by ___
remove, herbivory
Types of animal disturbance
gator holes (biodiverse), prairie potholes, muskrats (eat outs), beavers (alter water flow and levels)
Important disturbances in praire potholes
grazing, mowing, and natural disturbance
Peat-cutting
cutting out peat from a watland to enhance plant biodiversity and peat is used as fuel, however must continually remove peat
Logging
created by removing biomass from a forested system, however secondary effects are greater (ruts, roads, and canals)
Logging can also have …
permanent hydrological changes
Hurriance disturbance has 4 main components
felling of trees (wind), saltwater pulses (storm surge), freshwater pulses (rain), and sediment redistribution (waves)
Frosts can convert ____ to ___
mangrove swamp, salt marshes
Patches in wetlands are formed by …
fires, prolonged flooding, muskrat herbivory, ice scour, floating mats of litter, buried alluvial deposits
Invertebrates are ___ to disturbances and when they are at ___ densities there is ___ food for waterfowl
senstive, high, more
What are the disturbances that cause high invertebrate densities (highest to lowest)
cutting cattails → buring → mowing → flooding
Ways to measure and determine effects of disturbances
Measure relative changes in species compostition and biomass
Determine how the effects of disturbance varies between habitats or groups of species
Competition
negative effects that one organism has on another, examples consumption and resource limitation
Competition takes ____ studies over a ___ period of time
multiple, long
Ghost of competition past
species that use to overlap (over a resource) divereged characteristics to partion resources and aviod competition; past competition shapes current morphology and behavior
Competition is often _____ causing ___ competition, one species ___ another
one-sided, asymmetric, out competes
Assymetric competition is the greatest and least in which habitats
greatest: fertile and flooded
least: unfertilized and moist
Competition for ____ produces ____ hierarchies through processes such as
light, competitive, rates of vegetative reproduction, antagonism (competition), and dense canopies suggest that the competition is for light
Prescence of canopy has ___ effects on growth
negative
____ plants are often larger than ____ plants
Dominant, subordinate
Survival of the weak competitor can be due to
patches, disturbance, and gradients

Centrifugal organization
links high competition with low diversity, with the ocre habitats as fertile, undisturbed, and have high biomass
Peripheral habitats are at risk due to
stablized hydrology, high amounts of nutrients, lowered herbivory, fire suppression, and genetic changes (including hybridization)
Natural disturbance regimes help these species …
bog turtles, box turtles, spotted turtles, wood turtles, baltimore checkerspot butterflies, bog buck moths, sedge wrens, and rare sedges and orchids
Allogenic burial
sediment and OM from external areas (deltas)
Autogentic
OM produced locally (peat bogs)
Sediment arrives in ___ and may not remain in the same location
single pulses
Sediment loads
increase with rainfall and deforestation AND produces a diverse array of wetland types AND decrease when dams are constructed
Natural levees
built by the river as it deposits new layers of sediment along the banks
Artifical levees (embankments)
Concrete walls put up to stop flooding
Rain-fed bog formation
peat accumulates across a broad area with no lateral expansion, peat can begin at multiple individual points and eventually fuse together, or peat could accumulate at one site and increase in depth and area
Burial may happen from ___ and reduces _____ and ___ . It can cover ___ and growing points. Controls which species survive and drives _____ in plant communities.
flooding, runoff, or storms; light and oxygen; shoots; changes
Rhizomes
store energy, allow lateral spread, push shoots through sediment
Which wetlands are most and least sensitive to burial
Most senstive: alpine
Tolerant: freshwater and coastal wetlands
How does burial depth affect plants
survival decreases as depth increases, deeper burial prevents emergence, and can lead to plant death
cm of sedi is sufficient to reduce emergence by more than how much? 2cm burial?
50%, near 0 emergence
Seedlings are senstive to burial because
burial reduces species diversity, sediment may contain contaminants, and only tolerant species persist
3 leading threats ot freshwater aquatic ecosystems
sedimentation, exotic species, impoundments
with 20% fish are extinct of declining
Primary sources for alter sediment loads and it consquences
agricultural non-point pollution and road construction, aquatice invert and fish eggs are suffocated and nutrients in sedi causes summer growth then after winterkill the O levels are reduces
When urban land increases ther should be an ___ in ____ to compensate
increase, riparian
Sediment cores indicate changes in vegetation in what pattern
open water → floating-leaved plants → reeds → bog
Counter succession factors
fire, flooding, erosion, burial, and climate change
Short term effects of burial
immediate death