-In mature cells, 90% of volume may be taken up by central vacuoles
-Bounded by vacuolar membranes (tonoplasts)
-Filled with cell-sap (water fluid, slightly to moderately acidic)
-Contains dissolved substances (salts, sugars, organic acids, small proteins)
-Frequently contains anthocyanins (red, blue, purple water-soluble pigments)
-Functions: maintenance of cell pressure and pH, storage of numerous cell metabolites and waste products