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artificial selection
deliberate manipulation by humans, as in animal and plant breeding, of the genetic composition of a population by allowing only individuals with desirable traits reproduce
natural selection
the process by which individuals with certain heritable traits tend to produce more surviving offspring than do individuals without those traits, often leading to a change in makeup of the population
1) heritable variation leads to 2) differential reproductive success
evolution by natural selection occurs when...
fitness
the ability of an individual to produce viable offspring relative to others of the same species
adaptation
any heritable trait that increases the fitness of an individual with that trait, compared with individuals without that trait, in a particular environment
natural selection, genetic drift, gene flow, mutation
what are the four processes that change allele frequencies?
hardy-weinberg principle
a principle of population genetics stating that genotype frequencies in a large population do not change from generation to generation in the absence of evolutionary processes (e.g. mutation, gene flow, selection) and non random mating
gene pool
all of the alleles of all the genes in a certain population
1) random mating 2) no natural selection 3) no genetic drift (random allele frequency changes) 4) no gene flow 5) no mutation
what are the five assumptions of the hardy weinberg principle?
inbreeding
mating between closely related individuals; increases homozygosity of a population and often leads to a decline in the average fitness via selection (inbreeding depression)
inbreeding depression
in inbred offspring, fitness declines due to deleterious recessive alleles that are homozygous, thus exposed to selection
no, b/c non random mating only changes genotype frequencies, not allele frequencies
is non random mating an evolutionary process?
deleterious
in genetics, referring to any mutation, allele, or trait that reduces an individual's fitness
sexual selection
a type of natural selection that favors individuals with traits that increase their ability to obtain mates; acts more strongly on males than females
genetic variation
the number and relative frequency of alleles present in a particular population; also the proportion of phenotypic variation in a trait that is due to genetic rather than environmental influences in a certain population in a certain environment
directional selection
a mode of natural selection that favors one extreme phenotype with the result that the average phenotype of a population changes in one direction; usually reduces genetic variation in a population
purifying selection
selection that lowers the frequency of or even eliminates deleterious alleles
stabilizing selection
a mode of natural selection that favors phenotypes near the middle of the range of phenotypic variation; reduces overall genetic variation in a population
disruptive selection
a mode of natural selection that favors extreme phenotypes at both ends of the range of phenotypic variation; maintains overall genetic variation in a population
balancing selection
a mode of natural selection in which no single allele is favored in all populations of a species at all times; instead there is a balance among alleles in terms of frequency and fitness
heterozygote advantage
a pattern of natural selection that favors heterozygous individuals compared with homozygotes; maintains genetic variation in a population, thus is a form of balancing selection
frequency dependent selection
a pattern of selection in which certain alleles are favored only when they are rare; a form of balancing selection
INTERsexual selection
the sexual selection of an individual of one gender for mating by an individual of the other gender (usually by female choice)
INTRAsexual selection
competition among members of one gender for an opportunity to mate (usually male-male competition)
sexual dimorphism
any trait that differs between males and females
ecological/environmental selection
a type of natural selection that favors individuals with heritable traits that enhance their ability to survive and reproduce in a certain physical and/or biological environment, excluding their ability to obtain a mate
genetic drift
any change in allele frequencies due to random events; causes allele frequencies to drift up and down randomly over time, and eventually can lead to the fixation or loss of alleles
sampling error
the selection of a non-representative sample from some larger population, due to chance
founder effect
a change in allele frequencies that often occurs when a new population is established from a small group of individuals due to sampling error
genetic bottleneck
a reduction allele diversity resulting from a sudden reduction in the size of a large population due to a random event
gene flow
the movement of alleles between populations; occurs when individuals leave one population, join another, and breed
mutation
where do entirely new alleles come from?
point mutation
a mutation affecting only one or very few nucleotides in a gene sequence
lateral gene transfer
the transmission of genes from the parental generation to offspring via sexual or asexual reproduction
beneficial allele
an allele that allows individuals to produce more offspring
fossil record
all of the fossils that have been found anywhere on Earth and that have been formally described in the scientific literature
population thinking
the ability to analyze trait frequencies, event probabilities, and other attributes of populations of molecules, cells or organisms
extant species
a species that is living today
transitional feature
a trait that is intermediate between a condition observed in ancestral (older) species and the condition observed in derived (younger) species
vestigial trait
a reduced or incompletely developed structure that has no function or reduced function, but is clearly similar to functioning organs or structures in closely related species
phylogenetic tree
a branching diagram that depicts the evolutionary relationships among species or other taxa
homology
similarity among organisms of different species due to their inheritance from a common ancestor: DNA sequences, proteins, body parts
genetic homology
similarity in DNA nucleotide sequences, RNA nucleotide sequences, or amino acid sequences due to inheritance from a common ancestor
developmental homology
a similarity in embryonic form, or in the fate of embryonic tissues, that is due to inheritance from a common ancestor
structural homology
similarities in adult organismal structures that are due to inheritance from a common ancestor
morphology
the shape and appearance of an organism's body and its component parts
if mutation, selection, and genetic drift cause isolated populations to diverge sufficiently; genetic isolation and genetic divergence
what causes speciation to occur?
lack of gene flow
what causes genetic isolation?
selection, genetic drift, and mutation in isolated populations
what causes genetic divergence?
speciation
the evolution of two or more distinct species from a single ancestral species
species
an evolutionarily independent population or group of populations; generally distinct from other species in appearance, behavior, habitat, ecology, genetic characteristics, etc.
the biological species concept
the definition of a species as a population or group of populations that are reproductively isolated from other groups. Members of a species have the potential to interbreed in nature to produce viable, fertile offspring but cannot interbreed successfully with members of other species
1) the biological species concept 2) the morphospecies concept 3) the phylogenetic species concept
what are the three criteria used to identify a species?
pre-zygotic isolation
reproductive isolation resulting from any one of several mechanisms that prevent individuals of two different species from mating
post-zygotic isolation
reproductive isolation resulting from mechanisms that operate after mating of individuals of two different species occurs; the most common mechanisms are the death of hybrid embryos or reduced fitness of hybrids
morphospecies concept
the definition of a species as a population or group of populations that have measurably different anatomical features from other groups
polymorphic species
a species that has two or more distinct phenotypes in the same interbreeding population at the same time
cryptic species
a species that cannot be distinguished from similar species by easily identifiable morphological traits
phylogenetic species concept
the definition of a species as the smallest monophyletic group in a phylogenetic tree
monophyletic species/clade/lineage
an evolutionary unit that includes an ancestral population and all of its descendants but no others
synapomorphy
a shared, derived trait found in two or more taxa that is present in their most recent common ancestor but is missing in more distant ancestors; useful for inferring evolutionary relationships
subspecies
a population that has distinctive traits and some genetic differences relative to other populations of the same species but that is not distinct enough to be classified as a separate species
allopatry
condition in which two or more populations live in different geographic areas
allopatric speciation
speciation that occurs when populations of the same species become geographically isolated, often due to dispersal or vicariance
vicariance
the physical splitting of a population into smaller, isolated populations by a geographic barrier
sympatry
condition in which two or more populations live in the same geographic area, or close enough to permit interbreeding
sympatric speciation
the divergence of populations living within the same geographic area into different species as the result of their genetic (not physical) isolation
sister species
closely related species that occupy adjacent branches in a phylogenetic tree
polyploidy
the state of having more than two full sets of chromosomes, either from the same species (autopolyploidy) or from different species (allopolyploidy)
autopolyploid
the state of having more than two full sets of chromosomes due to a mutation that doubled the chromosome number; all the chromosomes come from the same species
allopolyploid
the state of having more than two full sets of chromosomes due to hybridization between different species
reinforcement
the natural selection for traits that prevent interbreeding between recently diverged species
hybrid zone
a geographic area in where interbreeding occurs between two species, sometimes producing fertile hybrid offspring
phylogeny
the evolutionary history of a group of organisms
tree of life
the most universal of all phylogenetic trees, depicting the evolutionary relationships among all living organisms on earth
tip
the end of the branch on a phylogenetic tree that represents a specific species or larger taxon that has not yet produced descendants-- either a group living today or a group that ended in extinction
outgroup
a taxon that is closely related to a particular monophyletic group but is not part of it
ancestral trait
a trait found in the ancestors of a particular group
derived trait
a trait that is clearly homologous with a trait found in an ancestor of a particular group, but that has a new form
parsimony
the logical principle that the most likely explanation of a phenomenon is the most economical or simplest; in relation to phylogenetic trees, it suggests that the one requiring the fewest evolutionary changes is most likely to be correct
homoplasy
similarity among organisms of different species due to reasons other than common ancestry, such as convergent evolution (ex: wings of birds and bats)
convergent evolution
the independent evolution of similar traits in distantly related organisms due to adaptation to similar environments and a similar way of life
evo devo
evolutionary developmental biology, focuses on how changes in developmentally important genes have led to the evolution of new phenotypes
adaptive radiation
rapid evolutionary diversification within one lineage, producing many descendant species with a wide range of adaptive forms
fauna
all the animal species characteristic of a particular region, period, or environment
basal metabolic rate
the total energy consumption by an organism at rest in a comfortable environment
ficks law of diffusion
a mathematical relationship that describes the rates of diffusion of gases