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Prognathism
Protrusion of the lower jaw or mandible
Occipital
Back of the skull
Thorax
chest, pleural cavity
Evolution
Change over time; in a biological sense refers to how the human body has gradually changed over generations
Species
A group of organisms that can interbreed and produce fertile offspring
Ex. homo sapiens (humans)
Adaptation
A biological response, typically a change, of an organism to its environment
Ex. bipedalism
Human Osteology / Skeletal Biology
Study of skeletal structure, function, and variation in humans
Anthropometry
Measure of anatomical dimensions and variability
Bioarchaeology
Study of skeletal material from archaeological sites; study things like bones from burials to understand certain rituals or routines regarding death in past civilizations
Paleopathology
Study of disease and trauma in past populations
Forensic Anthropology
Applied anthropological subfield using osteology and archaeology for legal applications
Human Population Biology / Variation
Study of how humans vary in response to their environment, specifically stressors, in a physical way
Ex. difference in altitude, climate, etc.
Molecular Anthropology
Study of genetics of modern humans, non-human primates, and human ancestors
Primatology
Study of the behaviours and biology of non-human primates
Ex. diet, communication, behaviours, etc.
Paleoanthropology
Study of the evolution of humans, ancestors / fossil relatives, including the fossil primate record, to understand the circumstances leading to modern humans and primates
Genus
Grouping of different species based on shared similarities
Strata
Layers of rock
Taxonomy
Science of naming / classifying organisms
Catastrophism
Explanation for extinction, theory that earth's landscape is the result of catastrophic events
Development of Evolutionary Thought
Pre-16th Century: fixity of species + great chain of being / scala naturae
16-17th Century: Heliocentrism (Galileo + Copernicus), laws of physics + gravity, scientists open to idea of evolutionary theories
Darwin: coined the concept of natural selection, Wallace invented similar concept + collaborated w Darwin
Mutation
Change in sequence of chemical bases
Can also occur in response to environmental conditions or replication error
Source of new population in a population
Must occur in a gamete (sex cell) to be evolutionary
Modern Evolutionary Theory
A change in allele frequency from one generation to another
A two step process
Production and distribution of variation
Natural selection acting on variation
Gene flow
Interchange of genes between populations
Individuals mate in new population, but don't necessarily stay there
Genetic Drift
Random, occurs in small populations
Alleles become more / less prevalent
Biological Species Concept
A species is a group of interbreeding or potentially interbreeding organisms that produce fertile offspring that are reproductively isolated from all other such groups
Extant vs Extinct
Extant means species still exists / is alive
Extinct means species is not still living and reproducing
Homology
Similarities between organisms due to descent from common ancestor
(e.g., tetrapod, forelimb, bone number + form)
Primate Characteristics
Flexible, generalized limb structure; affords various degrees of flexibility and allows mammals to engage in multiple forms of locomotion
Prehensile (grasping) ands and feet, sometimes tails; differs from most mammals
Pentadactyl; five digits on hands and feet
(e.g., fingers, toes)
Quadrupedal, moving on four limbs
Bipedal, moving on two limbs
Arboreal, active in trees
(e.g., feeding, sleeping, socializing)
Teeth variation dictated by species' habitats / food availability; each tooth type serves diff purpose
Strepsirrhines
Most primitive primates
Lemurs, Madagascar
Lorises, South Asia + Africa
Galagos, Africa
Fossils
Preserved remains, impression, or trace of an organism that once lived
Organic parts of organism are replaced by inorganic materials
Some parts perverse better than others
Trace fossils: Impression of an organism
(e.g., footprints)
Dating Methods
Relative dating: older / younger than
Chronometric / absolute dating: used to determine actual age
Isotopes: variations of a chemical element
Radiometric decay: measure of rate of decay in isotopes; occurs at predictable and measurable rate
Potassium-argon (K/Ar) dating
The Geological Timescale (Cezonic Era)
Holocene, post ice age(0.01)
Pleistocene, ice age (2.6-0.1)
Pliocene, modernization of mammals (5.3-2.6)
Miocene, increase in temp + animal population (23-5.3)
Oligocene, plains + deserts + grasslands become common (34-23)
Eocene, first neanderthals (55-34)
Paleocene, extinction of dinosaurs (65-55)
Early Hominin Characteristics
Bipedalism, contrast from other primates
Increase in brain size
Manufacture and use of tools
Evolution of Primates
During the paleocene, plesiadapiforms were the earliest form of primates that existed and lived, however they lacked some of the characteristics that we define primates by today
Hominin Origins
Last common ancestor (LCA) shared by chimpanzees and humans lived appx 6-9 million years ago (late miocene) in Africa
Hallmark hominin characteristics have been linked to changing environmental changes
Mosaic Evolution
When characteristics evolve at different rates; combo of primitive + derived traits
Early Homo Characteristics
Pleistocene epoch
Adapted in response to environment
Homo Habilis
East + South Africa, 2.4-1.4 Ma
Stone tools
Short, small teeth
Larger brain (640 cc)
Orthognatic, smaller teeth
3.5 - 4.5' in height, 70 lbs average
Stone Tools
Indicates material culture
Simple, deliberately manufactured
Homo Erectus
1.8 Ma - 100 000 years ago
Larger brains, smaller dentition
Africa, Asia, and Europe
Modern body proportions
Reduced sexual dimorphism
Larger brain (900 cc), smaller dentition
Modern body proportions
Zhoukoudian HE = from China
Acheulean Industrial Complex
Africa, 1.7 Ma - 160 000 years
Handaxes, cleavers, bifacial tools
More standardized / complex than oldowan
Oldowan Tools
Used by Homo habilis
Oldest known stone tools, 2.5 Ma
Controlled Use of Fire
1.5 - 1.0 Ma, debated
800 000 - 400 000 years, good / widespread evidence
Social + survival benefit
Archaic Homo
~700 000 - 200 000 years
Africa, Europe, Asia
Derived + primitive characteristics
Large brain (1200 cc)
Midfacial prognathism, protecting occipital
Controlled fire (790 000 years)
Natural shelters + spears (400 000 years)
Wooden spears (400 000 years)
Homo Neanderthalis
200 000 - 28 000 years
Europe, west Asia, Middle East
Used fire, shelter, clothes, tools
Low + elongated cranium
Midfacial prognathism
No chin
Occipital bun, heavy brow ridge
Large brain size (1200 - 1700 cc)
5'1 - 1'5 in height, 1190143 lbs in weight
Fossil Record Trauma Evidence
Low life expectancy
Evidence of injury + healing
Characteristics of Mousterian Tool Technology
Flake tools
Soft hammer percussion
Use of adhesives
When was the Neanderthal Extinction + Why did it happen?
Within 10 000 years of modern humans arriving (42 000 years)
Possibility of modern humans having a competitive advantage
Denisova Cave
195 000, 76 000 - 52 000 years
Best known from DNA
Genetic similarities to melanesians, australian aborigines, philippines, southeast asians
Homo floresiensis
100 000 - 60 000 years
Liang Bua Cave, Flores, Indonesia
Small, long, low cranium (417 cc)
Reduced prognathism
No chin, mandible like early homo / australopiths
Long arms + feet, short legs
Just over 1m in stature
Early Views of Neanderthals
Neander Valley (1856)
La Chapelle-aux-Saints, France (60 000 years)
Severe osteoarthritis
Enamel Hypoplasia
Growth arrest lines
Anatomically Modern Homo Sapiens
Africa, 315 000 years
Evolved from archaic
Large brain size, globular cranial vault
Gracile mandible + chin
African Origin Model
Homo sapiens evolved in africa, expanded ranges without interacting w/ archaic groups in Europe + Asia
Multiregionalism
Homo sapiens evolved simultaneously in Africa, Europe, Asia; gene flow maintained single species + regional variation
Assimilation Model
Homo sapiens have african origins + interbreeding
Humans Discovered: Jebel Irhoud, Morocco
Homo sapiens discovered in 1961, originally dated to 160 000 years; actually 315 000 years
Ice-Free Corridor Model
Low sea levels during Pleistocene
Archaeological sites predate ice free corridors
Coastal Route Model
Early Americans expanded ranges from Asia, followed Pacific coast of the Americas southward 17 000 - 15 000 years ago
Supported by rich coastal ecosystems
Projectile Point
Used as arrow / spear points
Omo Kibish, Ethiopia (104 000 years)
Barbed Harpoon Point
Katanda, DR Congo (80 000 - 90 000 years)
Burin
Used for engraving + carving
La Madeleine, France (17 000 - 11 500 years)
Awls
Used for piercing small holes in leather, wood, etc.
Blombos Cave, South Africa (77 000 years)
Bone Sewing Needles
Xiaogushan, Liaoning Province, China (30 000 - 23 000 years)
Parietal Art
Cave paintings, pigments came from minerals
Earliest paintings found in Sulawesi, Indonesia, 45 500
Endogamy
Mating within a distinct group, causes similar populations
The Neolithic Evolution
The neolithic (or new stone age) was 11,700 years ago
Large distribution of humans while staying as our own original species (humans), due to adaptability skills
Changes in settlement patterns, new technology, biological repercussions
Sedentism → permanent settlements, communities, cities
Domestication and the Origins of Agriculture
Breeding due to selected traits often prevents animals' survival in the wild
Animals considered domesticated when they cannot breed / survive without human intervention
Plants considered domesticated if they genetically differentiate from their wild relatives
Sedentism
Settlement pattern of staying in one place, work to maintain domesticated species
Civilization
Stage of human social and cultural development and organization that is complex
Cities
An urban centre that both supports and is supported by a hinterland of lesser communities
e.g., Çatalhöyük, Uruk
City Characteristics: Çatalhöyük, Turkey
Complex settlement with 8000 residents
2000 houses, multi-level w access via roof
9000 years ago
City Characteristics: Uruk, Iraq
"First true city"
5500 - 1800 years ago
Sumerian civilization, 20 000 people
Temples at society core
Included government, military, social classes
Civilization Characteristics: Mesopotamia (Iraq)
Uruk, 5500 - 1800 years ago
Ebaid, early farming culture, 7500 - 6200 years
Drainage systems → agricultural productivity → population growth
Sumerian was first urban civilization in the region of Sumer, 4900 - 4350 years ago; true social stratification
Ziggurat
Mud-brick temple-pyramids build on elevated platforms
Civilization Characteristics: Indus Valley
3600 - 2900 years ago
Architectural planning: irrigation, retaining walls, platforms, sewer systems
Civilization Characteristics: China
Longshan period
4600 - 4000 years ago
distinct social classes
Ritualistic
Warfare
walled towns
Shang dynasty
3600 years ago
First historic civilization of china
Elite ruling class
Writing system + divination
New World Civilizations
Relies on agriculture + plant domesticates
Long distance trade
Social classes + powerful leaders
Low reliance on wheel
Civilization Characteristics: Mesoamerica
Olmec
3200 - 2400 years ago
Sites at San Lorenzo and La Venta
Significant landscape modification
Ritualistic sculpture and carving (anthropomorphic), courtyards, plazas, artificial ponds
Written communication w glyphs
Maya
2100 years ago
Trade networks
Complex tools
Class + religion based society
Writing system w hieroglyphs + codices
Organized sport