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1869 Fredrich Miescher — his proposal for DNA
studied pus cells
believed that nuclein (not protein) is the genetic material
1900s Phoebus Levene — his proposal for DNA
renamed nuclein to DNA/RNA
discovered nucleotides (sugar, phosphate, and 1 of 4 bases)
proposed nucleic acids have a tetranucleotide structure (repeating sequence of 4 nucleotides), TOO SIMPLE
tetranucleotide arrangement made the world shift back to protein idea
1928 Frederick Griffith
injects rats with r-strain (rough) & s-strain (smooth = virulent)
r-strain - mice lived, s-strain - died, heat killed s-strain - lived, killed s-strain+ living r-strain - died
transforming principle — something from the heat-killed bacteria transformed living bacteria (non-pathogenic) to become deadly
1944 Oswald Avery, Macyln McCarty, Colin MacLeod
continued with pneumonia
tested for protein, nuclein & DNA
destroyed DNA and unable to do transforming principle (proved DNA is used in transformation principle)
not ready to publish
1952 Alfred Hershey, Martha Chase
blender experiment — radiolabeled viral proteins (s-35) & viral DNA (p-32) and injected into bacteriophage
DNA has no sulfur, proteins has sulfur, no phosphorus (very clever experiment)
proved DNA had genetic material
1949 Erwin Chargaff
DNA structure — % composition and base pair rules (A + T, C + G)
1953 Rosalind Franklin
3D DNA structure —> X-Ray diffraction
double helix w/ repeating pattern every 3.4nm & 0.34nm
1954 Watson & Crick
used Chargaff’s rule and Franklin’s findings
shows H-bonds, double helix, turn clockwise every 10 nucleotides, 5’ — 3’ direction, antiparallel
how DNA is wrapped EUKARYOTES
wrapped around histones (proteins) for protection & reduce volume
nucleosome — group of 8 histones
chromatin — nucleosome arranged in loops (scaffold)
in mitosis — chromosomes (visible)
how DNA is wrapped PROKARYOTES
DNA resides in nucleoid — no nucleus
eubacterial chromosomes are found in loops that join together, making a ring, can supercoil
could have plasmids (accessory loops)
archaea have circular chromosomes associated with histones