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Who were the primates of Paleocene?
Plesiadapiformes 65-56 MyA
Who were the first primate of eocene
Adapoids - early streph
omomyoids. -early tarsier or haph
56-34 MyA
First Haphlorrhines my have
May have evolved in the Eocene, but radiated during Oligocene (34-23 MyA)
First Homonoids (Apes)
22-5 MyA
Where did oldest Apes go?
Late/mid Miocene, climate cooled with less tropical rainforests, reduction of ape taxa.
Pliocene-Pleisocene
Last 5 Mya
Reduced forests, increased radiation of monkeys, decreased in hominoids
Modern Synthesis of Evolutionary Theory
Adds Malthusian competition, variation, mutation, medelian genes and genetic inheritance to the idea of natural selection
Extended Evolutionary Synthesis Adds
Plasticity, Inclusive inheritance, and Niche Construction
Niches are
context for evolution
includes the structural, temporal, and social context in which a species exists. It includes space, structure, climate, nutrients, and other physical and social factors
Plasticity/Accomodatin
Developmental, or phenotypic, plasticity is the capacity of an organism to change its phenotype in response to the environment.
Accommodation: the mutual and often functional adjustment of parts of an organism during development that typically does not involve genetic mutation
Inclusive Inheritance
Inheritance occurs beyond just the genetic level – epigenetic changes can be inherited
Multiple pathways of Inheritance
genetic, epigenetic, behavioral and symbolic/cultural) can affect
evolutionary processes
Characteristics of Hominin evolution
Bipedalism
Brain size increase
Tool Use
Fire and Food
Culture
What happened in Micoene?
Micoene 23-5 Mya; temperatures drop, more seasonal, more wood and grasslands
many species, died, changed, or lost numbers
Why Bipedalism
Feeding adaptations in Miocene apes
harvesting fruit from woodland trees
Keeping cool
hands free for other things
How are human different from apes?
Habitual bipeds
wide and paraboilic dentition
thick molar enamals,
large molars
small canines and no CP3
long juvenile period and slow development
big brain and culture through social language
LCA of Chimp and humans
Diverges around 7-5 Mya, still unknown
Sahelenthropus Tchadensis
5-7 Mya in Chad, no CP3, small brain and maybe bipedal with browridge
Orrorin Tugensis
6 Mya in Kenya, maybe bipedal, climing lmbs and teetj were a mix of chimp and human
ardipithecus Ramidus
4.4 Mya in Ethiopia
Bipedal, somoe prognathism with small brain
no cp3
opposable big toe and equal arm/legs
The Austrolopithcenes
Genus includes multiple species dating from 4.2 million to 1.4 million years ago
Mix of bipedal locomotion and ape-like features
Small-brained creatures (but bigger than chimpanzees) with lots
of sexual dimorphism in body mass and a little in canine size
Small incisors/canines relative to body size
Diet: herbivores & possibly some meat-eating
Some species had specialized dietary adaptations
Au. Anamensis
East Africa 4.3-3.9 MyA
Bipedal long forearms with ape fingers
large canines, prognathic and lives in wood/grasslands
Au. Afarensis
East Africa 3.6-3 Mya
Lucy (bipedal, but short wide pelvis)
small brain, prognathic
canines, large range of habitats
long femoral neck
Au. Africanus
3-22 Mya in South Africa
bigger brain than others. less prognathic, with small front teeth
shorter less dimorphic canines
central Foramen magnum
chimp like dentition
lots of c4 foods like tubers, grass, roots, seeds
Au. Garhi
Ethipia, 2.5 Mya
Sagittal crest, more. robustt
Bigger teeth
maybe used tools????
Au Sediba
South Africa .977 Mya
420 cc brain
small cheek teeth like humans
human like pelvis and hands
ape like feet
different ways of bipedalism
like au. africanus
What are the robust Australopithecines
Unlike Gracile forms, around 2.5 Mya a new more robust form known as paranthropus evolve. special diet and different jaw/teeth
Robust Au’s have:
Large cheek teeth
huge chewing muscles and broad flared cheek bones
large temporal fossa
post orbital constriction
P. Aesthiopicus
Ethiopia, 2.5 Mya
small brain and prognathic with otherwise robust features
P. Bosei
East African, 2.2-1.3 MyA
Hyper Robust- sexually dimorphic body size
4-5 hundred cc
sagittal crest, heavy chewer
P. Roustus
South African form
1.8 1 mya
500-530 cc brain
sagital crest chewer
see root nuts and c4 plants
Kenyanthropus Platypos
only one species? is it a varient of au?
3.5-3.2 ya
less prognathic but small brain
only face found
Early Homo
2.3 Mya East Africa Larger brain and more human teeth
The Pleistocene
2.5M to 12k years ago
ice ae but not all ice → glaciall then ccold/dry climate
fast cycling climate
climate slowed down around 1.25 MyA
creates new feeding niches
What are the agreed upon features of Homo
larger brain
smaller flatter face
small jaws and teeth
increased reliance on culture and stoneools to exploit more ofmthe environment
Other, less agreed on features of homo
Larger body size
longlegs short arms
reduce sex dimorph.
chin in later species
Ledi Geraru
2.8 Mya East Africa
could be earliest homo fossil
small dental and mandible
no chin
bipedal spine and homo teeth
Homo Habilis
oldest most widely accepted homo
2.3-1.5 Mya found in Tanzania
presumed to make tools
larger brain than Au.
no sagittal crest and small teeth
Homo Habilis, cont.
Found in East Africa with lots of variability
Post Cranial:
long strong thumbs, straight short fingers with broad fingertips
means they had more manual dexterity
H. Habilis Taxonomy
Ppl say that it might be two species
H. Hablis smaller brain and less robust
H. Rudolfensis. morerobust and big brain 2.4-1.6 mya
Olduwan tool industry
Mode One very simple stone tools
H. Ergaster
Africa 1.8-600k ya
East/South
Oldowanm tools
have eye narrowing eceeded forehead and no chin
but also less prgnathic, taller skull, small jaws, post canine teehs and projecting nose
larger brain
and limbs are similar to modern human
less dimorphic and have terrestrial life history
possible slow developmen
Homo Ergaster Tools
1.6-1.4 M Years ago they started using Mode 2 Acheulean tools alongside mode 1
more specific design
bifacial
hand axe→ cleavers and larger than modern 1 tools
H. Ergaster Diet
Most likely ate meat and had controlled use of fire
temprerate habitat
Use of cooking
Softens food
• Reduces toxins
• Reduces physical barriers
• Bursts cells
• Changes molecular structure
• Increases starch/fiber digestion (12-
75%)
• Increases protein digestion (6-35%)
• Increases dietary breadth
• Decreases cost of digestio
Leaving Africa
1.8 Mya H ergaster leaves africa for eurasia maybe becomes H Erectus
How are the new fossils different from early homo and Australopithecines
fossils of 800k to 300k YA have Diversified materials left with humans
large brain and body
less sex diffeences
shorter arms and longer legs
reduced teeth size
The Early Middle Pleistocene
900/300k YA
Colder more varied climate
migration to Eurasia
Climae changes shaped homo lieae
Ergaster became smarter
Homo Heidelbergensis
Afroeurasia
900/130k YA
Large rain with some primative features like low skull,, thick cranium, robust
but also had sidewalls and high foreheads and round skull backs
used refined tools (1 and 2)
Late middle/late pliestocene
300k in Africa mode 3 tools appear
Hom Floesiensis
The hobbit
14-16k YA
1 meter tall small brain and very primitive feet shoulder pelvis and waist
used flake tools
coexisted in Indonesia
Homo Naledi
300-236k YA
South Africa
Small heads and low EQ
small and thin
use of trees
burials and fire
Neanderthals
130-30k YA
big brain
short muscle body
low forehead big browridge
big face big nose
small teeh and worn out
occipital bun
personal adornments
burials and mode 3 tools
difficult lives
Early Homo Sapien
Africa 190 Kya omo ethiopia
large brain no specialization
large face Chin prominent brows
high rounded braincase
sophisticated tech and behaviour
Muddle In the Middle
3 theories of H sapien evolution and phylogeny
Multiregional theory
wolpoff- all one species all became humans
partial replacement
rightmire
we evolved in africa
went from ergaster to heidelburg to human/neanderthal
Complete replacemnt
evolved in africa completely replacing erectus than ergaster then neanderthals and humans
3 distinct, genetially isolated species
done :P