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Phylogenetic Trees
how to make traits used in reconstructing phylogenies
A branching diagram that represents a hypothesis about the evolutionary history of a group of organisms.
Uses of phylogenetic trees
Organizing knowledge of biological diversity
Allowing structured classification of organisms
Providing insight into evolutionary events
Demonstrate that living species are the result of
their evolutionary history
Show that all living species are contemporaries-
each organism is the product of parents that
successfully gave rise to offspring for the last >3
billion years
How traits evolved
biogeographic patterns
age of divergence
bioactive chemical searches
Phylogeny
history of the descent of a group of organisms from their common ancestor
Phylogenetic / Evolutionary Tree
shows order or pattern of when different species or groups of species evolved
When species evolve via "descent with modification" the
closer the species are together the more characteristics should be (mores common characters the closer related)
Systematics uses
fossil, molecular, and genetic data to infer evolutionary relationships
(living organism)
How are phylogenetic trees constructed?
trees constructed with analysis of characters or traits
Characters can be
morphological, molecular, or behavioral
Apomorphic
Derived characters (those possessed by some taxa on the tree but not all and not the common ancestor)
Plesiomorphic
Ancestorial characteristic (ancestors had this trait and descendants have it as well)
Homologus
similar due to common ancestor (good)
Analogus
misleading due to convergence (bad)
Homologous Characteristics
inherited from a common ancestor, hence due to common ancestry (sometimes has different functions though)
Bird vs. Insect Wing
analogous
Bird vs. Bat Wings
Homologous
Analogus Characters
NOT due to inheritance but similar due to convergence/ parallel evolution
Adaption for a similar function
Analogus
Vultures same or different?
New World - from storks
Old World - from raptors
Convergence doesn't come from same species but look similar because adapted to same environment
ex. Sharks, Dolphin, Ichthyosaur
Divergent Evolution
evolution of one or more closely related species into different species; resulting from adaptations to different environmental conditions
Divergent evolution may make homologues traits appear different
ex. vertebrate limbs (common origin, but vastly different)
Tenants
Species change over time
species are related by common ancestor
Morphological evolution
Whales and Hippos
What traits are used to reconstruct phylogenies
nucleic acid sequences and molecular data
When did humans and apes slit
7 - 10 mya
Parsimony
simplest explanation of evolution is the best explanation (fewest evolution characters)
Bayesian analysis
to revise and update initial assessments of the event probabilities generated by the alternative solutions. (makes more readable)
maximum likelihood analysis
likelihood is the probability of the data given a tree, its branch lengths and a model of evolution
Embryo development evolution
human embryos go through all former common ancestors (tail, gills, notochord, Chordata, etc.) before final form
Phenetics (numerical taxonomy)
uses both apomorphines and pleomorphisms (groups based on overall similarity)
Evolutionary phylogenetics
uses apomorphines, but recognizes monophyletic and non-monophyletic groups in classification
Cladistics
uses only apomorphias, but recognizes only monophyletic groups in classification
Classification systems
improve ability to explain organism relationships (reflects evolutionary history, memory aid, and unique universally used names)
Monophyletic tax tonic groupings
should be used in our classification systems
monophyletic group
a common ancestor and all of its descendants (reflect evolutionary history in classification)
Molecular clocks
the average rate at which a species' genome accumulates mutations, used to measure their evolutionary divergence and in other calculations.
Phylogenetic trees help with
classification, disease control, crop development, conservation
Polyphyletic
pertaining to a group of taxa that includes distantly related organisms but does not include their most recent common ancestor
paraphyletic
Pertaining to a group of taxa that consists of a common ancestor and some, but not all, of its descendants.