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Messy Phylogeny
The phylogeny appears messy, but color schemes and lineage traces reveal phylogenetic signals.
Horizontal Transfer
Associated with endosymbiosis.
Mitochondria and Chloroplasts
Key endosymbiotic features.
Endosymbiosis
Multiple rounds involve primary, secondary, and sometimes tertiary stages.
Eight Major Lineages
Alveolates, Straminopiles, Rizarians, Excavates, Plants, Amoebozoans, Fungi, and Animals.
Multicellularity
A key innovation in eukaryotic diversity.
Choanoflagellate
The closest relative to all animals.
Alternation of Generations
Land plants alternate between sporophyte and gametophyte stages.
Alveoli
Air sacs under the cell membranes
Pseudopods
Extensions of the cytoplasm with cytoskeletal elements used to move and feed
Carotenoids
Photosynthetic pigments in chloroplasts
Mycelium
The main body of the fungus
Gametangia
Organs forming gametes that are protected
Synapomorphy
Plant embryo is protected
Waxy Cuticle
Prevent water loss
Closable Stomata
Regulate gas exchange and water loss
Globophytes
The earliest branching lineage after the first endosymbiotic event
Major Groups
There are three major groups with horizontal transfers between them.
Acquisition Order
Mitochondria were acquired first, followed by chloroplasts.
Chloroplast Formation
Photosynthetic bacteria were taken up to form chloroplasts.
Mitochondria Membranes
Eukaryotes with mitochondria typically have two membranes (inner and outer).
Pigment Variation
Different pigments mean different bacteria were taken up.
Primary Endosymbiosis
Primary endosymbiosis involves a non-photosynthetic eukaryotic cell engulfing a cyanobacterium.
Peptidoglycan Remnants
Peptidoglycan remnants suggest the evolutionary story.
Secondary Endosymbiosis Definition
Other photosynthetic eukaryotes, including some protists, have undergone secondary endosymbiosis.
Membrane Count
Usually, three membranes are seen, but sometimes four.
Flexible Cell Membrane
A flexible cell membrane allows the cell to move and engulf other cells.
Diversification Period
These diversified during the Precambrian period (1.5 billion to 600 million years ago).
Impact of Multicellularity
It allowed animals to become larger and more complex.
Artificial Selection Experiments
They used unicellular organisms and selected for traits that favor multicellularity.
Choanocytes Resemblance
Choanocytes, found in sponges, resemble choanoflagellates.
Dinoflagellates Abundance
Dinoflagellates are abundant and contribute to organic debris on the ocean floor.
Coral-Dinoflagellate Relation
Corals use dinoflagellates for carbohydrate production.
Straminopiles Flagella
They have two flagella, but some species have reduced or lost them.
Diatoms Silica Deposition
They deposit silica within structures resembling cell walls, creating different shapes.
Diatomaceous Earth Use
Diatomaceous earth contains diatom shells with sharp, jagged edges, used to prevent pest invasions.
Brown Algae Pigments
They have carotenoids, giving them a brownish color.
Foraminifera Limestone
They create sedimentary deposits that compress into limestone.
Radiolarians Endoskeleton
They secrete a glassy endoskeleton.
Cellular Slime Molds Vegetative State
In a vegetative state, they are single-celled with a haploid nucleus, reproducing through mitosis and fission.
Opisthokonts Flagella
If flagella are present, they are on the back (posterior) of the cell.
Limestone Production
Rizarians (specifically, Foraminifera) are responsible for much of the limestone production due to their calcium carbonate shells.
Saprobes Definition
Saprobes absorb nutrients from dead organic matter.
Hyphae Definition
Hyphae are tubular branches that extend immense distances underground.
Dry Rot Definition
Fungal infections can encroach within the xylem of plants, causing dry rot.
Chloroplasts as Synapomorphy
Chloroplasts are a synapomorphy (shared derived trait).
Waxy Cuticle Definition
Waxy cuticle to prevent water loss
Mycorrhizae Benefits
Beneficial associations with fungi (mycorrhizae) to promote nutrient uptake
Early Plant Lineages
The earlier lineages of plants were nonvascular and needed to live close to water sources.
Gamete and Spore Production
Gametes are produced by mitosis, and spores are produced by meiosis.
Early Land Plants
Early land plants (bryophytes) had a large gametophyte body.
Sporangium Function
The tip of the sporangium ruptures, releasing haploid spores into the environment.
Sporophyte Growth
Which means what follows fertilization is a sporophyte which grows out of the gametophyte.