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What are the 5 characteristics of life?
1) Organisms are made of cells
(STRUCTURE & FUNCTION)
2) Organisms are capable of replication
(INFORMATION FLOW, EXCHANGE AND STORAGE)
3) Organisms have mechanisms to store information
(INFORMATION FLOW, EXCHANGE AND STORAGE)
4) Organisms have a mechanism to acquire & transform energy
(PATHWAYS & TRANSFORMATION OF ENERGY &
MATTER)
5) Organisms have a capacity to adapt/change
(EVOLUTION)
What is the modern cell theory?
All organisms are made of cells and all cells come from pre-existing cells
Scientist Spotlight: Al-Jahiz
Abū ʿUthman ʿAmr ibn Baḥr
al-Kinānī al-Baṣrī
• 776 - 868
• He was reported to haveat
written over two hundred
works
• Wrote “Book of Animals”
• Grouped animals
• Transformation of animals
• Effect of environmental
factors
Who were the 3 people that developed the thought of Evolution, and what did each one contribute?
1) Plato (428-348 BC)
• Typological
Thinking
• Special
Creation
• Unchanging
2) Aristotle (384-322 BC)
• Great Chain of
Being
• Hierarchy
• Unchanging
3) Jean Baptiste de Lamarck (1744-1829)
• Species change over
time...
• Up the great chain of
being
• Mechanism:
Inheritance of
acquired
characteristics
What did Darwin and Wallace contribution to Evolution?
Populations evolve overtime and are linked together through common ancestry; not structured into a hierarchy.
Population definition
Population: a group of individuals of the same species
that are living in the same place at the same time
than can and do breed
Differing views of evolutionary mechanisms between Lamark and Darwin (giraffes)
Lamarck : giraffes lengthened necks by
stretching
• Mechanism: acquired characteristic passed to offspring.
• Darwin: long-neck giraffes out compete short
necks
• Short neck giraffes died or don’t breed

Theory of Natural Selection?
1) Common ancestry: Species today
arose from preexisting species.
2) Descent with Modification: Species
characteristics can change over time
3) Natural Selection: The process whereby organisms better adapted to their environment tend to survive and produce more offspring.
How old is the Earth?
Earth is about 4.6 billion years old
Where and when did the earliest signs of life appear?
Earliest signs of life appears in rocks about
3.4-3.8 billion years ago
What are the 3 ways in how species change throughout time?
1) Extinction
2) Transitional Forms
3) Vestigial Traits
What is Extinction?
Termination of an organism meaning reproduction is no longer able to happen.
Evolution also supports natural selection by creating opportunities for new species to emerge and thrive. When a species faces extinction, it leaves behind ecological niches that become available to other organisms. This vacancy creates evolutionary opportunities for surviving species to diversify and adapt to fill these roles, often leading to adaptive radiation events where one lineage rapidly diversifies into many new species.
Who was Baron Georges Curvier and what did he discover about extinction?
Baron Georges Cuvier (1769-1832):
1) groundbreaking paleontologist who compared living organisms with fossils (among other accomplishments)
2) Realized two extinct species (Irish Elk and Helicoprion) were too large to escape notice, and too unique to be an existing species.
How do species change through transitional forms?
Living species “succeed” fossil species in the same region. → refers to the idea that as species evolve, they leave behind their remains, and that new species are reconstructed through the fossil remains.
Vestigial Traits definition
Vestigial Traits – reduced or incompletely developed structure that has no or reduced function but is like functioning organ or structured in closely related species.
Two examples of vestigial traits?

Describe the process of speciation
The formation of new species due to reproductive isolation or environmental changes; splitting of an ancestral species into two or more distinct species
Speciation occurs when populations of a species become reproductively isolated, preventing gene flow and leading to genetic divergence over time.
Homology definition
“Study of likeness” or similarities between organisms that is due to inheritance from a common ancestor.
What are the 3 different types of homology?
1) Structural Homology
2) Developmental Homology
3) Genetic Homology
What is structural homology?
1) refers to similarities in anatomical structures
2) also knows as comparative methodology: compare physical structures in different organisms to identify common ancestry.
3) this similarity can be observed in the bones, organs, and genes that evolved from a common ancestor.
What is developmental homology?
1) similarities in developmental patterns; studying embryonic development patterns to illustrate evolutionary patterns.
2) embryos of many vertebrae species develop in similar ways.
3) all vertebrates go through a stage in which they have four limb buds, a tail, and a series of gills.
What is genetic homology?
1) similarities in DNA sequences
2) a concept in genetics that describes the relationship between genes or DNA sequences that are similar because they have been inherited from a common ancestor.
What does evolution favor?
evolution favors traits that enhance survival and reproductive success, which may include simple traits that are more efficient.
What is biological fitness?
Biological fitness is the ability of an individual to produce surviving offspring.
How is fitness determined?
by observing the traits of an organism that are more advantageous in regard to responding to the demands of the environment.
What is an individual?
A single organism
What is a population?
A group of individuals that are the same species at the same time at the same place.
Natural selection acts on the ___, while evolution occurs on the ____ level.
individuals, population
1) individuals with traits that enhance survival and reproduction pass these traits to offspring, leading to gradual changes in populations overtime.
What is Darwin’s four postulates?
1) Variation exists among individual organisms that make up populations.
2) Some of the trait differences are heritable
3) Survival and reproductive success are highly variable. (many more offspring are produced than can possibly survive meaning offspring has to compete for resources for survival; not random)
4) The subset of individuals that survive best and produce the most offspring is not a random sample of the population.
In modern biology we condense Darwin’s four postulates into a simple sentence:
Evolution occurs by natural selection when heritable variation leads to differential reproductive success.
Dr. Zuleyma Tang-Matinez
1) Born in Venezuela
2) 1st Latin America President of the International Animal Behavior society
3) 1st Hispanic women to receive tenure at University of Missouri
4) Research focuses on social structure in mammals and reptiles, including how it impacts population genetics
5) Has challenged Bateman principles (sexual)
Population genetics
studies change in allele frequency and genotypes in populations over time.
The field was spurred by the combination of Mendelian genetics with Darwinian evolution (known as Modern synthesis).
How do you find determine allele frequency?
Allele frequency = # times allele appears in population/ total # alleles in population
Gene pool
all possible gametes (egg and sperm) from each generation in a single gene pool.
What is the mathematics behind Hardy-Weinburg?
Frequency of A (p) + Frequency a (q) = 1
What are the 3 possible genotypes?
1) AA (p²)
2) Aa (2pq)
3) aa (q²)
The Hardy-Weinberg Principle states
The frequencies of both alleles in a population will remain constant from generation to generation.
The Hardy-Weinberg Principle has __ assumptions and what are they?
1) Mating is random
2) No natural selection
3) No genetic drift
4) No gene flow
5) No mutation
The Hardy Weinberg is important as a concept because it serves as the mathematical ____ to study evolutionary processes.
null hypothesis
What are the 2 main frameworks of Natural selection?
1) Ecological or environmental selection
2) Sexual selection
Directional Selection
1) Favors one extreme phenotype over others, shifting the population's traits in one direction.
2) Selective pressures favor individuals with traits at one end of the phenotypic spectrum, favoring the favored phenotype. overtime, the traits that are more advantageous become more common, while the less advantageous diminish in frequency.
3) decreases genetic diversity; can remove deleterious alleles from population which is called purifying selection
Stabilizing Selection
1) Favors intermediate phenotypes, reducing variation in a trait.
2) natural selection is more likely to favor the average or most common trait within a population, because the intermediate phenotype is most likely to be fit for that current environment.
3) Overtime this can result in a population with reduced genetic diversity as the selection process removes the more severe phenotypes.
Disruptive Selection
1) Favors extreme phenotypes at both ends of the spectrum, potentially leading to speciation.
2) can lead to speciation by creating a scenario were populations become increasingly adapted to different environments which can cause barriers to arise, ultimately resulting in the emergence of new species.
Balancing selection
1) No single allele has a distinct advantage
2) Maintains genetic variation
3) One type favors rare alleles (frequency-dependent selection)
Sexual Selection
Selection based on traits that enhance mating success rather than survival.
Genetic Drift definition
Random changes in allele frequencies in a small population due to chance.
key point: genetic drift is random with respect to fitness.
What are some examples of how genetic drift can lead to significant changes in a small population?
1) In the founder effect, a small group of individuals breaks away from a larger population to establish a new population. Because this new population is formed from a limited number of individuals, the allele frequencies in the new population may differ significantly from those in the original population.
2) The bottleneck effect occurs when a population experiences a dramatic reduction in size due to environmental events (like natural disasters) or human activities (such as habitat destruction). This reduction can lead to a loss of genetic diversity, as only a small number of individuals contribute to the gene pool of the surviving population. The alleles present in the surviving individuals may not represent the original population's genetic diversity, leading to significant changes in allele frequencies.
Gene Flow definition
The transfer of alleles or genes from one population to another.
1) Occurs when individuals migrate between populations and interbreed, they bring different genetic material, which can enhance the overall genetic variation.
2) causes allele frequencies to be more alike between the two populations
Natural Selection definition
Differential survival and reproduction of individuals based on advantageous traits.
Mutation definition
A change in DNA that can create new alleles in a population.
1) They introduce genetic variation, which is essential for a population's adaptability to changing environments.
2) This variation can lead to new phenotypes, some of which may confer advantages in survival and reproduction.
3) Random with respect to fitness; usually deleterious or neutral
4) Happens “slowly”, except for organisms with short generation times
What are the 3 ways mutations can happen?
1) point mutation: a single or small number of base pairs changed in DNA
2) chromosome-level mutation: change in the number of structures of chromosomes
3) lateral gene transfer; movement of genes from one species to another
Sexual Selection definition
A form of natural selection where individuals with certain traits are more likely to attract mates.
1) Cause competition for mates
2) Changes frequency of alleles and increases fitness
What are Darwin’s contributions to the ideas of sexual selection?
concluded that male peacock tails must be adaptive by attracting mates and increasing reproductive success.
Nonrandom mating
Occurs when there is interbreeding or mate choice leading to changes in genotypes in a population but not allele frequency.
Why is interbreeding harmful?
Interbreeding leads to a loss of fitness overtime
Increased homozygosity: Inbreeding increases the likelihood of offspring inheriting two copies of the same harmful recessive allele, leading to inbreeding depression.
Reduced genetic diversity: The loss of genetic diversity within a population can reduce its ability to adapt to new environmental challenges, making it more susceptible to extinction.
Intersexual selection
a form of sexual selection where one sex chooses mates of the opposite sex based on specific traits.
Intrasexual selection
competition between members of the same sex and drives traits that are useful for direct combat
What hypothesis is Sexual Selection based on?
Bateman-Trivers Hypothesis
What is the pattern, process, and prediction of the Bateman-Trivers Hypothesis?
Pattern: males generally benefit more from multiple matings than females do, leading to greater sexual selection pressure on males to compete for mates.
Process: based on how much or how little a male or female invests in their offspring which determines how picky or non-picky they are with mating.
Predictions: any allele that increases a male’s attractiveness to females or success in male-to-male competition should rapidly increase in the population.
What does sexual selection lead to?
Sexual dimorphism
Sexual dimorphism
the differences in appearance between males and females of the same species, which can include variations in color, structure, size, to attract mates.
Joseph L. graves Jr. (1955)
1) First Black Scientist to receive Ph.D. in Evolutionary Biology in 1988
2) Evolution as a specialty did not form until 1946
Speciation results from
genetic isolation and genetic divergence
How does genetic isolation happen?
results from lack of gene flow between populations which prevent equalizing population allele frequencies
How does genetic divergence happen?
divergence results from mutation, genetic drift, and natural selection which proceeds independently in isolated populations
Species definition
a distinct, identifiable population(s) that is thought to be evolutionarily independent of other populations.
usually distinct by appearance, behavior, habitat, ecology, and genetic characteristics
What are the 3 ways in how species are defined?
1) Biological Species Concept
2) Morphospecies Concept
3) Phylogenetic Concept
Explain the Biological species concept
1) Talks about reproductive isolation; if no reproduction happens, no gene flow can occur
2) Pre-zygotic: prevents fertilization between different species)
3) Post-zygotic: prevents formation of reproductive zygote; hybrid offspring cannot survive or reproduce
What are the 5 mechanisms of reproductive isolation in pre-zygotic isolation?

What are the 2 mechanisms of reproductive isolation in post-zygotic isolation?

What are the advantages and disadvantages of in the biological species content?
1) advantages: intuitively appealing
2) disadvantages:
cannot evaluate reproductive isolation in fossils or asexual species
or in living species with ranges that do not overlap
What is the Morphospecies Concept?
refers to defining a species based on shared physical characteristics; states if 2 organisms loom alike, they are the same species, but if they differ significantly, they are diff species.
believe that differences arise because populations were independent and isolated
What does the Morphospecies Concept not account for and explain each one?
1) Cryptic species: groups of organisms that are not morphologically indistinguishable from one another but are genetically distinct enough to be considered separate species.
2) Polymorphic species: a species that exhibits two or more distinct forms within the same species.
e.g= peacock
through color, size, shape, and behavior
What are the advantages and disadvantages of the morphospecies concept?
advantages:
Useful when RIMs cannot be
applied
Widely applicable (fossils,
asexual, sexual species
disadvantages:
Highly contentious
Subjective
What is the phylogenetic concept?
The phylogenetic concept refers to the study of evolutionary relationships among all forms of life on Earth, an ancestral population and all its descendants
What are the advantages and disadvantages of the phylogenetic concept?
advantages:
Widely applicable
Logical: species form with
low flow & isolation)
disadvantages:
phylogenies not widely
available
Reflect a larger number of species
than other concepts?
In speciation what are the two ways species can form?
1) Allopatric speciation (different geographical location)
2) Sympatric speciation (same geographical location)
What is allopatric speciation type 1?
Allopatric Speciation 1: Isolation by Dispersal
Colonization is important for
speciation
1. Physical separation eliminates
Gene Flow
2. Genetic Drift, mutation and
Natural Selection causes
divergence.

What is allopatric speciation type 2?
Allopatric Speciation 2: Isolation by Vicariance
Physical Separation is important for
speciation
1. Physical separation eliminates
Gene Flow
2. Natural Selection, Drift and
Mutation causes divergence

What is Sympatric speciation?
refers to same geographical location
external (extrinsic events) = disruptive selection
internal (intrinsic events) = polyploidization
What is the process of sympatric speciation?

What is sympatric speciation 1?
disruptive selection (external)
different populations use different ecological niches meaning different populations within an ecosystem occupy various specific roles and positions they play within their environments. These niches are influenced by both biotic and abiotic factors, such as food availability, predators, temperature, and soil nutrients. Each population's niche is shaped by its unique set of conditions and interactions, allowing it to thrive and contribute to the overall health of the ecosystem.

What is sympatric speciation 2?
polyploidization (internal)
refers to autopolyploid and allopolyploid
poly: many
ploidy: # of complete chromosome sets in a cell
autopolyploid: an organism that has more than 2 complete sets of chromosomes, from same species.
allopolyploid: an organism with multiple sets of chromosomes derived from different species, typically formed after hybridization between 2 species.

What happens when isolated populations come
into contact?
Possible outcomes following secondary conduct

What is population thinking?
Population thinking emphasizes the variation within populations, leading to an understanding that evolution is a change in the genetic makeup of populations over time.