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arthopoda
arthropods
subphylum chelicerata (water mites & spiders)
subphylum pancrustacea (crustaceans & hexapods)
types of metamorphosis
ametabolous
hemimetabolous
holometabolous
simple metamorphosis
ametabolous
no change in organism between stages, except for size
incomplete metamorphosis
hemimetabolous
immature insects look similar to adults. Nymphs don’t have wings and adults have wings
complete metamorphosis
holometabolous
immature look nothing like adults
hexapods
6 legged
three body segments (head, thorax & abdomen)
hexapod orders
collembola
ephemeroptera
odonata
plecoptera
hemiptera
coleoptera
neuroptera & megaloptera
trichoptera
lepidoptera
diptera
collembola
glue-peg; springtail
minute hexapods
with or without jumping structure (furcula)
up to 8 ommatidia
short antennae
mostly terrestrial, few aquatic species
ametabolous
ephemeroptera
bioindicators
EPT taxa
short-lived wings; mayflies
wings held above body at rest
2 or 3 cerci
leafy gills on abdomen
only order where they molt again after having functional wings
hemimetabolous
found in lentic or lotic water
larvae feed on a variety of food
adults do not feed, they reproduce and die
odonata
tooth; dragonflies & damselflies
bioindicators
compound eyes
3 ocelli
antennae short & bristle-like
suborder: zygoptera (damselflies)
suborder: anisoptera (dragonflies)
carnivores/ predators as larvae & adults
adult stages are easy to ID, larvae pretty much all look the same
hemimetabolous
plecoptera
folded-wing; stoneflies
EPT taxa
associated with clean, cool running water
somewhat flattened, soft-bodied
poor fliers
aquatic larvae
hemimetabolous
many emerge during winter months
heimptera
half-wings; true bugs
large & diverse group
wings are part membranous, part leathery
hemimetabolous
coleoptera
sheath-wings; beetles
largest order of insects (+28,00 NA species)
forewings modified into wing covers (elytra)
hard & leathery, usually meet in a straight line on the back
holometabolous
neuroptera & megaloptera
alderflies, dobsonflies, fishflies, hellgrammites
originally grouped together
larvae are mostly predaceous
adults vary
holometabolous
neuroptera
nerve-wings; lacewings, antlions & owlflies
soft-bodied
four membranous wings
many cross veins & extra branches
some larvae are aquatic
adults are weak fliers
holometabolous
megaloptera
alderflies, dobsonflies and fishflies
bioindicators
hellgrammites (dobsonfly larvae) are the largest stream insects, well adapted for hunting in riffles
life cycle:
females lay their eggs on trees that overhang streams
young falls into the water and lives in the stream for three years
emerges and pupates on land and turns into the adult
adults are usually nocturnal
trichoptera
hair-wings; caddisflies
EPT
somewhat mothlike appearance
4 membranous wings that are hairy
larvae: collectors, some predators
build cases and collect food, have hook-like appendages on abdomen
holometabolous
lepidoptera
scale-wings; butterflies & moths
scales on wings
large compound eyes
holometabolous
diptera
two-wings; true flies
forewings present, hind wings absent
hind wings modified into halteres (maintains equilibrium while flying)
holometabolous
not good bioindicators
crustaceans
paraphyletic group
cladocerans and copepods are key primary consumers in many lakes & ponds
crustacean orders
ostracoda
copepoda
branchipoda
decapoda
isopoda
amphipoda
ostracoda
seed shrimp
shells are important in paleolimnology
occur in most habitats, even hypersaline and hot springs
copepoda
important zooplankton in both freshwater and marine systems
larval stage- nauplius
eggs can be resistant to drying
branchiopoda
cladocera (water fleas), fairy, tadpole, brine, and clam shrimp
primarily in lotic environments
cladocerans can produce eggs encased in an ephippium than can remain dormant for centuries
decapoda
crayfish, crabs, shrimp
mostly marine or estuarine, but many freshwater species
isopoda
terrestrial pill bugs
many marine species, and few freshwater species
generally detritivores
amphipoda
scuds and sideswimmers
similar in size to isopods, but laterally flattened
omnivorous and scavengers
why are macroinverts good bio-indicators?
taxa are differentially sensitive to pollution
they are not very mobile and thus cannot escape short- or long-term pollution events
they are relatively long lived
they are present in most streams, even those that lack fish
relatively easy to catch, view, and identify