the process in which two species undergo reciprocal evolutionary change through natural selection
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Qualitative natures of species interactions can...
Be altered if the background environment changes
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Niche
range of physical and chemical conditions under which that species can survive and produce (ex: hydra living is a specific temperature, salinity, and pH)
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N-dimentional hypervolume
species that coexist differ in some aspect of their niche
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Fundamental role
The environmental conditions under which a species can survive and reproduce; sometimes called a physiological niche; the set of environmental conditions under which a species can persist
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Realized niche
Portion of the fundamental niche that a species actually uses as a result of interactions with other species
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Competition can restrict the...
Fundamental niche
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What can restrict the realized niche?
-Presence of predators or pathogens -mutualism/commensalism
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Alpha = 1
Intraspecific and interspecific are equal to each other
- simplest and best known two species model of competition - provides foundation for many other ecological models - predicts a full range of outcomes, depending on the values of K's and alpha's
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Competition exclusion
One species eliminates the other
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Stable coexistence
Persist together forever
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Unstable coexistence
Persist together until perturbed
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Interspecific competition affects...
The populations of 2 or more species adversely (-/-)
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Intraspecific and Interspecific competition are likely occurring
Simultaneously
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2 forms of Interspecific competition
Exploitation and interference
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Most types of Interspecific interactions can be classified as 1/6 types
Individuals of one species inhibit individuals of another by consuming a shared resource (squirrels, birds, etc eating acorns)
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Preemption competition
Individuals of one species prevent occupation of an area by individuals of another species (sessile organism such as barnacles and clams)
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character displacement
increased ecological differences between species in regions where they occur together
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ecological release
the expansion of a species niche under conditions where their competitor species is absent; niche of the competitively-inferior species expands in the absence of the competitively-superior species
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observational competition studies
- negative correlations between species - attributed to present or past competition ("ghost of competition past") - cant determine cause and effect - other factors may be involved
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experimental competition studies
- addition/removal studies - manipulate presence and/or density of would-be competitors - must account for density effects - provides strong interference (strong evidence for or against) - difficult to do for many species
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Connell, 1961
- determined factors regulating distribution of Cthamalus stellatus and Semibalanus balanoides - one of the first studies to show interspecific competition through manipulative experiments
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fundamental niche depends on...
physical (abiotic) conditions
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realized niche depends on...
biotic and abiotic conditions
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competitive exclusion principle
states that complete competitors cannot coexist
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complete competitors
two distinct species that live in the same place and have exactly the same ecological requirements
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competitive exclusion requires that...
- competitors require exactly the same resources - environmental conditions remain constant
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competition is influenced by
non-resource factors
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environmental features that are not resources can...
influence the outcome of competition between species (ex: trout species)
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temporal variation in the environment...
influences competitive interactions
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why are there so many species?
- non-resource variability - resources varying in time - resources can be a variety of things, water, light, food, microhabitats - disturbance
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predation
consumption of one living organisms by another
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predators are
heterotrophs
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carnivores
consume animal tissue
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herbivores
consume plant or algal tissue
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omnivores
consume plant and animal tissue
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true predator
kills its prey immediately upon capture, more of less
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predators consume multiple prey organisms and function as ____________ throughout their lifetimes
agents of mortality
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lethal effects
- predators directly affect mortality rates through total combustion - prey directly affect predator birth/death rates - form modeled by L-V predator/prey model
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predators may_______ affect prey birth/mortality rates through partial consumption (herbivory or parasitism)
directly
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predators may __________ affect pre birth/ mortality rates through effects on prey behavior
indirectly
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Lotka and Volterra on predation
- developed 2 linked equations, one for prey and one for predator - plot results on a phase plane
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Lotka-Volterra Predator- Prey Model
predicts population cycles
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high latitude animals often have
population cycles
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Optimal foraging: Type 1
- number of prey consumed is linearly related to prey population size - more prey available, more are eaten
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prey switching
predator doest eat the prey at low densities, relying on a different food source instead
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apparent competition
a shared predator can make it appear that two species compete when they dont; two species that do not compete directly for resources affect each other indirectly because they share the same predator
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competitive release
a predator can reverse competitive exclusion
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trophic cascade
a predator of an herbivore can help a plant (the enemy of my enemy is my friend)
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chemical defenses
Compounds released by prey to defend themselves from predators.
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prey weapons
prey having physical defense mechanisms like hard shells or thorns to protect or fight their predators
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plant defenses: physical and chemical
plants evolved spines, thorns, and chemical toxins, such as morphine, strychnine, and nicotine, against herbivores.
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feigning death
faking death
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predator swamping
a prey strategy in which the per capita predation rate is reduced at high prey density
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schooling, flocking, group living
prey that tend to group together in order to survive (ex: prairie dogs, birds)
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escape tactics
being fast and/or maneuverable
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camoflauge & startling displays
an adaptation that allows an organism to blend in with its envoronment
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symbiosis
the intimate and protracted association between two or more individuals of different species; can be positive, negative, or neutral
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all parasitic relationships are...
symbiotic relationships
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larger species is typically considered the...
host species
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parasitism
- feed on the live host organism - an intimate relationship, with the parasite living on or in the host at least part of its life cycle - actively is harmful but generally not lethal, at least in the short term
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parasitoids
- attack the prey indirectly by laying eggs on the host's body - and intimate association with a single host - the eggs hatch and the larvae feed on the host, eventually killing it
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parasites increase their fitness by using the host in a close, prolonged association for
food, habitat, & dispersal; usually dont kill the host
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Host fitness is often decreased by the parasite through
individuals of one species grow over individuals of other species, inhibiting access to a resource (taller plants shading shorter plants)
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chemical interaction competition
individuals of one species release growth inhibitors or toxins that inhibits or kills other species (allelopathy in plants- secretion of chemicals that inhibit germination of other species)
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territorial competition
behavior of one species that excludes another species from a specific location that is defended as a territory ( a bird keeping other birds from nesting in its territory)
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encounter competition
non-territorial encounters of individuals of different species affect one or more of the species involved (scavengers fighting over a dead animal carcass)