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polynesian-smino (manifesting) (15.1-15.3)
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special senses
convey specific stimuli from specialized sensory organs in discrete locations of the head. differ from general senses in stimuli detected, structure of sensory receptors, and location of sensory nerves
transduction
when a physical or chemical stimulus is converted into an action potential that can be interpreted by the brain. special senses do this with stimuli such as light, chemicals, or sounds in the environment
olfaction
sense of smell/detection of odorants (chemicals) in the air. bypasses the thalamus.
olfactory epithelium
small patch of specialized epithelium located in the superior region of each nasal cavity. contains olfactory neurons, basal cells, and supporting cells
olfactory neurons
modified bipolar neurons that detect odorants. also known as chemoreceptors. consist of an apical modified dendrite, a cell body, and a basal axon. the modified dendrite projects into the roof of the nasal cavity. neurons are bundled into groups of 10-100 and travel through tiny holes in the cribriform plate of ethmoid bone to reach CNS.
olfactory cilia
increases surface area available for odorant detection. extend into a layer of mucus produced by olfactory glands.
olfactory nerve
formed by bundled groups of axons from olfactory neurons. axons terminate by synapsing on other neurons called mitral cells
olfactory bulb
structure located in the brain just superior to the ethmoid bone and inferior to the frontal lobe of the brain.
basal cells of olfactory neurons
stem cells that develop into olfactory neurons. ensure continual replacement of these neurons.
supporting cells of olfactory neurons
columnar cells that support and surround the olfactory neurons. Contain a pigment that gives the olfactory epithelium its color
olfactory tract
axons that exit the olfactory bulb; travel to other regions of the CNS for interpretation
step 1 of activation of olfactory receptors
binding of an odorant to its receptor, which activates a G-protein
step 2 of activation of olfactory receptors
the activated G-protein triggers the enzyme adenylate cyclase to convert ATP into cAMP
step 3 of activation of olfactory receptors
cAMP opens ion channels that allow sodium and calcium ions to enter the cell
olfactory pathway
olfactory stimuli are transmitted to the CNS and delivered to various regions of the brain
step 1 of the olfactory pathway
The axons of olfactory neurons (making up the olfactory nerve) carry olfactory stimuli to the olfactory bulb in the CNS.
step 2 of the olfactory pathway
An olfactory stimulus travels from the olfactory bulb to the primary olfactory cortex in the temporal lobe.
primary olfactory cortex
located in the inferomedial temporal lobe near the optic chiasma, is responsible for awareness and identification of an odor.
taste buds
Clusters of receptor cells and supporting cells on the tongue that detect taste sensation. each is contacted by sensory neurons that transmit stimuli to the central nervous system. typically located on the lateral surfaces of the papillae.
papilae
Finger-like protrusions on the surface of the tongue, some of which contain taste buds.
vallate papillae
largest and are dome-shaped; each contains hundreds of taste buds
fungiform papillae
mushroom-shaped and each contains only a few taste buds.
foliate papillae
ridges on the sides of the tongue and contain taste buds only in childhood.
filiform papillae
long, thin cylinders scattered across the tongue. lack taste buds but have sensory nerve endings that detect the texture and temperature of food.
gustatory cells
specialized epithelial cells with microvilli that contain receptors detecting different tastes. microvilli are found on the apical surface of the cell and project into a small pocket on the surface of the papilla, called the taste pore. basal end of each cell forms a synapse with the receptive endings of a sensory neuron that carries taste stimuli into the CNS via the facial, glossopharyngeal, or vagus nerve.
basal cells of taste buds
stem cells that differentiate into new gustatory cells, which have a lifespan of only 10-14 days
supporting cells of taste buds
surround the gustatory cells. They provide physical support but have no role in taste sensation.
sweet taste
mostly due to simple sugars such as glucose and fructose
sour taste
produced by hydrogen ions, as found in the citric acid of lemon juice.
salty taste
presence of metal ions such as sodium and potassium ions.
bitter taste
imparted by many different compounds, including alkaloids such as those found in coffee, many plant poisons, and rancid foods. humans are most sensitive to this taste, which serves a protective function
umami taste
often associated with meat or broth, is produced by glutamate or other amino acids.
step 1 of activating a taste receptor
Changes in ion movements depolarize the gustatory cell’s plasma membrane
step 2 of activating a taste receptor
Depolarization of the membrane opens voltage-gated calcium ion channels, and calcium ions enter the cell.
step 3 of activating a taste receptor
The calcium ions trigger the release of neurotransmitters, which produce an action potential in the axon of the sensory neuron
step 1 of gustatory pathway
Axons of the facial, glossopharyngeal, and vagus nerves carry taste stimuli from the tongue into the CNS.
step 2 of gustatory pathway
Axons of these three nerves terminate in the solitary nucleus in the medulla oblongata by synapsing on central sensory neurons
step 3 of gustatory pathway
Axons from the solitary nucleus synapse on neurons in the thalamus, which then send the taste signals to the primary gustatory cortex in the parietal lobe.
primary gustatory cortex
integrates incoming stimuli, leading to awareness and identification of a particular taste.