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What are the major 5 species
1. Cow
2. Pig
3. Sheep
4. Goat
5. Horse
What are some of the barriers to domestication?
• Diet
• Growth Rate
• Issues with Captive Breeding
• Nasty Disposition
• Tendency to Panic
• Social Structure
What is the Asiatic Mouflon an ancestor of?
Sheep
What is the Bezoar an ancestor of?
Goat
What is the Aurochs an ancestor of?
Cattle
What is the Wild boar an ancestor of?
Pig
What are Wild horses an ancestor of?
Horses
What are the uses of large domestic animals?
• Meat
• Milk products
• Fertilizer
• Land transport
• Leather
• Military assault vehicles
• Plow traction
• Fiber (wool, hair)
What is a breed?
A group of domesticated animals sharing common ancestry and distinctive
characteristics (e.g., conformation, color, or performance ability) that are consistently
transmitted to their offspring because of selective breeding. These traits are generally
documented and maintained by a breed registry
What is a standardbred?
n the 18th and 19th centuries, animals recorded in a breed registry based on having reached a
specific standard
What is a purebred?
An animal whose parents and ancestors belong to the same recognized breed.
Their lineage is recorded in a closed studbook or registry.
What is a crossbred?
An animal resulting from the intentional mating of purebred parents from two
different breeds
What is a landrace?
A population of domesticated animals that has developed over time through natural
selection and traditional management within a specific local environment
Who is robert bakewell?
Pioneer of systematic animal breeding
• Carefully selected males and females to breed
• Advocated use of inbreeding to create lines
that bred truly
• Used progeny tests for performance-based
selection
• Developed the New Leicester sheep breed from
the Lincoln Longwool
Who is Sir John Sebright?
Theoretical influence on breed
development
• 1809 essay, "The Art of Improving Breeds of
Domestic Animals"
• Recognized that laws drove inheritance, but
it was not critical to know them to use
selection
• Selection of males and females to breed
together considering each other's merits and
defects
• Supported ancestry breeding over progeny
testing
Who is Gregor Mendel?
The father of modern genetics
1. Innately curious and observant
2. Mendel was the first to take a quantitative,
scientific approach to the study of heredity.
3. Began his studies using mice
4. Switched to studying peas from 1856-1863
5. Kept careful records of 7 different traits
What is partial inheritance?
• Used pure-breeding lines for specific traits
• White flowers could be produced by crossing
two purple flowers, if the purple flowers had
at least one white flowered parent.
• Evidence that the factor for white flowers has
not blended irreversibly with the factor for
purple flowers.
• In particulate inheritance, heredity is the
product of discrete factors that control
individual traits.
What is a locus?
A specific position along a chromosome
What is an allele?
An alternative variant of a locus that arises by mutation
What is a wild-type allele?
The most common allelic form in a natural population
What does homozygous mean?
If both alleles of a particular gene are identical
What does heterozygous mean?
If the alleles are different from each other
What is hemizygous?
If only one copy of a gene is present
What is a genotype?
The complete set of alleles in an individual
What is a phenotype?
The visible or detectable effects of alleles on the structure or function of an individual
What was Mendel's First Law
The Law of Equal Segregation
-Offspring inherit one allele from each parent for a
trait, and no allele is favored or has an advantage
over the others
-Diploid individuals inherit two alleles for each
trait, with one allele on each member of a pair of
homologous chromosomes
-During gamete formation, the two alleles at a
gene locus separate and segregate from each
other
-Each gamete has an equal probability of
containing either allele
What was Mendel's Third Law?
The Law of Dominance
-For the two alleles of a gene, the dominant
allele is always expressed because it masks
the recessive allele
-Recessive traits are only seen when both
alleles are recessive
What is a punnett square?
• Matrix to predict all possible genotypes of
offspring
• If dominance relationship known, can predict
phenotype
What is haplosufficiency?
the wild-type allele
What is haploinsufficiency?
a genetic condition where a single functional copy of a gene is not enough to maintain normal function, causing a dominant disease or mutant phenotype
What is the structure of Livestock Industries?
Elite seedstock
Multiplier seedstock
Commercial producer
Growing & feeding enterprises
Who was King James 1
King James I imported Arabian stallions around 1616
-Any horse bred to an Arabian was "thoroughly bred"
-Thoroughbreds were tested for stamina by racing them at
Newmarket
What are the rules for the general stud book?
• Founded by James Weatherby in 1791 who was the keeper of the Match Book
• Early volumes retrospectively recorded pedigrees back to the late 1600s from private records
• Remains the official record of Thoroughbred pedigrees owned by Weatherbys
• Horses can be traced by direct descent to the original foundation bloodstock of the breed
• Inclusion in the studbook requires that a horse descend entirely through registered parents
• Arabians continued to be registered in the GSB until the 1960s
• The Thoroughbred became a purebred breed and has been genetically closed since the 18th century
• The GSB describes the biological and legal definition of a Thoroughbred
• The Jersey test
What are the rules for the american stud book?
• First published in 1868 in the United States by Sanders Bruce
• In 1896 ownership transferred to The Jockey Club
• Registers Thoroughbreds foaled in North America or imported from approved foreign stud books
• Regulates naming, parentage verification, ownership, transfers, exports and imports
What are the British Beef Cattle breeds?
• Angus
• Hereford
• Polled Hereford
• Red Angus
• Shorthorn
What is the angus?
Most Annual Registrations in US
• Originates from Aberdeen-Angus counties in
Scotland
• 300,000+ registrations in 2025
• US Herd book first published in 1886 is now closed
• Black, naturally polled, moderate size
• Excellent marbling, early compositional maturity
• Excellent maternal breed, easy fleshing, good milk
production
• Early maturing sexually
How does melanogenesis connect to selection for coat color?
Melanogenesis = the process of making melanin (pigment), which determines coat color. Genetic variation in melanogenesis genes changes the amount, type, or distribution of pigment, allowing breeders to select animals for desired coat colors.--Genes control melanin → melanin controls coat color → selection favors desired gene combinations."
What is the hereford?
2nd Largest Annual Registration in US
• Originated in Herefordshire, England
• Major US breed from 1920s to 1980s
• Red with white face to behind the ears
• White on the throat, underline, legs, switch
• Moderate size, adaptable, good disposition
• Low to moderate milk, relatively early maturing
• Poorest marbling of the British breeds
• Polled Hereford breed is naturally hornless
• Closed registry
What is the red angus?
Registry started in 1954
• Red, naturally polled, moderate size
• Excellent marbling, early compositional
maturity
• Excellent maternal breed, easy fleshing, good
milk production
• Early maturing sexually
• Red Angus Association allows some grading
up
What is the shorthorn?
Originated in England
• First breed to have a registry
• Three major colors - red, white, roan
• Moderate size, good disposition
• Excellent milk production
• Most numerous in the US until 1920s
• Not significant today
• Has appendix registry which allows outside
genetics to be introduced
What is the south devon?
Originated in England
• First breed to have a registry
• Three major colors - red, white, roan
• Moderate size, good disposition
• Excellent milk production
• Most numerous in the US until 1920s
• Not significant today
• Has appendix registry which allows outside
genetics to be introduced
What is the difference of continental beef cattle to british beef cattle?
• Larger framed and faster-growing than British breeds
• Later maturing and produce less milk than British breeds
• Produce leaner and more muscular carcasses with less marbling
• Used as terminal sires in a two-breed or three-breed cross
What is the charolais?
Originated in Charolles and Nievre, France
• White cattle in the region since 878 A.D.
• Imported to Mexico by Pugibet after WWI
• King Ranch purchased 2 bulls in 1936
• Purebred Charolais 31/32 blood
• White, off-white to cream coloration
• Extremely fast growing, muscular breed
• Large framed with heavy birth weights
• Lean, muscular carcasses
• Progressive ataxia
What is the simmental?
Fifth in annual registrations
• Straw colored through light red to dark red
• US Simmental red or black
• Unpredictable white spotting
• Gentle disposition
• Large framed, large mature size
• Heaviest milking continental breed
• Fast growing in the feedlot
What is the gelbvieh?
German "yellow" cattle
• Breed started in 1850 in Franconian districts of Bavaria
• Stringent selection to repopulate herds post WWII
• AGA started in 1971 and has 45,000 active registered
cows
• Any color, horned or polled, heavy milker
• Fullblood registration 100% Gelbvieh
• Purebred registration 81.6% to 99.9% Gelbvieh
• Balancer cattle are registered hybrid seedstock with 25%
to 75% Gelbvieh and the balance Angus or Red Angus
• Southern balancer are at least 25% Gelbvieh and 6.25%
to 50% tropically adapted breed
What is the Braunvieh?
German for "brown cattle"
• Breeders' society formed in 1897
• Second most populous breed in Switzerland
• Primarily used for dairy in Europe
• 130 animals imported from 1869 to 1880
became nucleus of American Brown Swiss
• Flourished as a beef breed in Mexico
WHat is the chianina?
One of the Oldest Breeds
• Originated in Italy and have different size and
type in different regions
• Largest type was US foundation stock in 1971
• Extremely large framed, late maturing, fast
growing breed
• Lean, muscular carcasses, suspect marbling
• Fullblood, purebreds, and composites
• Fullbloods from white to steel gray in color
• Short horns
• Skin color must be back for fullblood and
Chiangus animals
What is the limousin?
Third in annual registrations
• Originally golden red
• No color requirements in US registry
• Moderate sized continental breed
• Below average maternally, lacking in milk
• Extremely lean, muscular carcasses
• Fastest growing breed in the US
What is the salers?
Native to the Auvergne region of France
• Considered one of the oldest European breeds
• First imports into US in 1975
• DNA screening for beta-mannosidosis since 1995
• Excellent maternal breed
• Good carcass yield and carcass quality
• 28,000+ registrations per year
• Fullblood, purebred, and percentage Salers
• Optimizers have at least 6.25% but less than 75%
Salers blood
What is the Tarentaise?
Originated in the Tarentaise Valley in
the French Alps
• Breed named in 1859 and herdbook founded
in 1888
• First Tarentaise bull imported into North
America in 1972
• Developed in geographic isolation
• Adaptable to harsh environments
• Dual purpose breed
• Moderate size with good muscling
• Fullblood Tarentaise are horned
What are characteristics of American breed beef cattle?
• Bos indicus influence
• Composite design with stabilized composition
• Purpose-bred for southern US environments
• Enhanced heat tolerance
• Improved maternal ability
• Hybrid vigor (heterosis)
• Docility and manageability
• Beef quality and growth performance
What is the brahman?
Developed from 3 Indian breeds
• First American breed
• Originate from Guzerat, Gir, and Nellore
• First importation in 1849
• Red and grey strains with black pigmentation
• Horned
• Large pendulous ears
• Highly developed sweat glands
• Noted for heat tolerance, disease and insect
resistance, and crossing ability
• Poor marbling and slow sexual maturity
What is the beefmaster?
Developed by Lasater Ranch, TX
• From systematic crossing of Hereford,
Shorthorn, and Brahman cattle
• Recognized by USDA as a new breed in 1954
• No color standards
• The Six Essentials - Weight, Conformation,
Milking Ability, Fertility, Hardiness,
Disposition
• Largest number of registrations (~40,000)
What is the brangus?
Developed to combine superior traits
• Crosses began at USDA in Jeanerette, LA in 1932
• 3/8 Brahman, 5/8 Angus
• Moderate size
• Solid black, no brindling, and polled or scurred
• Excellent mothering ability
• Early sexual maturity
• Heat tolerance and foraging ability
• Intermediate in carcass merit
• Red Brangus, Ultrablack, Ultrared, Optimizer
What is the santa gertrudis?
Developed by Kleberg on King Ranch
• 5/8 Shorthorn, 3/8 Brahman composite
• Trace back to a single sire, Monkey
• Short, slick hair
• Deep cherry red color, no black pigment
• Noted for maternal ability of females
• Overall hardiness
• Santa Cruz (50% Santa Gertrudis, 25%
Gelbvieh, 25% Red Angus)
What is meiosis?
cell division that makes gametes (sperm & eggs).
1 diploid cell → 4 haploid cells
Cuts chromosome number in half
Creates genetic variation
Has 2 divisions: Meiosis I & Meiosis II
What are the stages of meiosis 1?
PMAT
Prophase I → homologous chromosomes pair up & crossing over
Metaphase I → homologous pairs line up in the middle
Anaphase I → homologous chromosomes separate
Telophase I → 2 haploid cells form
What are the stages for meiosis 2?
PMAT
Prophase II → chromosomes condense
Metaphase II → chromosomes line up in the middle
Anaphase II → sister chromatids separate
Telophase II → 4 haploid cells form
What are the stages of prophases in meiosis 1?
Leptotene → chromosomes condense
Zygotene → homologous chromosomes pair up (synapsis)
Pachytene → crossing over occurs
Diplotene → homologs begin separating; chiasmata become visible
Diakinesis → chromosomes fully condense; nuclear envelope breaks down
Lazy Zebras Prefer Doing Dishes
What is the ideal dairy type?
• Stature
• Angularity
• Level rump
• Long and lean neck
• Milk veins
• Strong feet and legs
What is the holstein?
Developed in North Holland & Friesland
• First imported to Boston in 1852
• Dominant breed of dairy cattle worldwide
• Bred and strictly culled for efficient
production on grass
• Large frame with unparalleled milking ability
for fluid milk production
• Black and white
• Red and white Holstein recognized as a
separate breed
What is the brown swiss?
Declared a US breed in 1906
• Large framed breed known for its docility
• Coat ranges from light brown with gray to
very dark brown.
• Light muzzle with light dorsal stripe
What is the ayrshire?
First US imports in 1822
• Only native Scottish dairy breed
• Several breeds, including Teeswater stock, Dutch and
Channel island imports were crossed to form
foundation animals
• Dark red and white cattle but no registry restrictions
on color patterns
• Horns are a foot long and curve outwards, but calves
typically dehorned
• Excellent udder conformation, minimal foot and leg
problems
• Moderate buffer fat
• Average 305 day lactation is 14,534 lbs
• Efficient on pasture and hardy in cold weather
What is the Guernsey?
High butterfat, high protein milk
• Developed from Alderney cattle brought to Guernsey
in 960 A.D.
• Coat color varies from yellow to reddish-brown with
white patches
• Introduced into US in 1840 and American Guernsey
Cattle club formed in 1847
• Aggressive young sire selection program for continual
breed improvement
• Consume 20-30% less food per pound of milk than
larger dairy breeds
• Adaptable to warmer climates
What is the jersey?
Excellent breed for cheese making
• Small, feminine cow
• Adaptable to many climates, more heat tolerant
• Excellent grazers
• Very fertile and early maturing sexually
• Light tan to dark fawn with darker shading on
head and legs
• Highest percentage butter fat and significantly
more solids-not-fat than other breeds
What is the difference between qualitative and quantitative traits?
Qualitative = categories → distinct traits, usually 1/few genes Ex: black vs. red coat
Quantitative = quantity → measured/continuous, many genes Ex: weight, height, milk production
What assumptions are made when estimating the number of loci for a quantitative trait?
Alleles are additive
All loci contribute equally
Loci are unlinked
Original parents are homozygous for alternative alleles
How do you predict the number of loci from the number of phenotypes?
Number of phenotypes = 2N + 1
N = number of genes/loci contributing to the phenotype
Solve for N to estimate the effective number of loci
What is the probability of getting an extreme phenotype in an F₂ population?
Probability = (1/4)ᴺ
N = number of genes/loci
Applies to either extreme phenotype
How do you calculate the probability of specific outcomes from multiple loci?
For independent loci, multiply the probabilities together