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Principle of Allocation
Limited resources must be allocated between growth, survival, and reproduction (example: animals forage, breed, care for offspring. Plants allocate biomass and nutrients to different parts like root, stem, etc)
A Trade-Off
Resources invested in one function are not available for another (an adaptation or trait that improves fitness in one area comes at the cost of another area due to limited resources) Example: species could have more smaller or less bigger offspring
Survivorship Curves
Type I: most reach old age (humans)
Type II: some reach old age (squirrels)
Type III: very few reach old age (plants)
Fast-Slow Continuum
Fast species tend to be small, while slow species tend to be large (many exceptions exist)
Case Study: DDT
DDT was used as a insecticide in the 1900s, caused eggshell thickness issues for many bird species, especially bald eagles
Demography
Study of how a population changes over time
The BIDE Model
Nt+deltat=Nt+B+I-D-E
Nt=number of individuals in a population at time t
Birth, Immigration, Death, Emigration
Simplified Version (assuming closed population): Nt+deltat=Nt+B-D
Population Growth Assumption
every individual has an equal chance of reproducing and dying
dNt/dt = b-d = rNt
r=each individual’s net contribution to the population
Exponential Growth Model
Nt=N0 x e^rt
N0 = initial number of individuals
r = intrinsic growth rate
When r is positive, the population increases, when r is 0, the population does not change, when r is negative, the population decreases
Per Capita Population Growth Rate
(1/N)(dN/dt)
Rate of population growth divided by population size, measures average rate of population change for a single individual
Density Dependence
Positive: per-capita population growth rate increases
Negative: per-capita population growth rate deceases
0: Equilibrium (b=d)
Logistic Model
(1/N)(dN/dt) = r (1-(N/K))
r = constant
K = carrying capacity
Population comes to equilibrium when N=K
(1/N)(dN/dt) is greatest when N is near zero
Density Independent
Sometimes, N is limited by something else and fluctuates (ex. volcano eruption impacts forest tree populations)
Competition
Species A and B try to acquire the same resources
Intraspecific: competition between individuals of the same species
Interspecific: competition between different species
Predation
Predators reduce prey abundance, so prey defend physically, chemically, escape, mimicry (honest: appears unpalatable, is unpalatable; dishonest: appears unpalatable, is palatable), fight back
Herbivory
Eating plants (the plant might or might not die, so it is only predation sometimes)
Usually a negative impact, but sometimes it could be beneficial
A cost paid to support mutualism (seeds eaten by squirrels who also disperse them)
Regular disturbance that removed dead tissue and reduce disease
A signal that promotes growth
Mutualism
Both species benefit (ex. disperse seeds, pollinate flowers, defend against parasites, gather nutrients, etc)
Facilitation
One species benefits another, the second species is unspecified, also called mutualism for plants (ex. host plants facilitate other plants growing inside them)
These interaction could shift from competitive to mutualistic as elevation or other stress increases
Fungus and soybean is competition in low phosphorus, mutualistic in medium phosphorus levels, and fungus becomes parasitic in high levels
Commensalism
One species benefits, the other is unaffected (ex. remora and host, a zebra shark)
Indirect Interactions
Exploitation Competition: two predators eat the same prey
Indirect Mutualism: plant A and B help each other when C is most palatable
Human Interactions
Agriculture, livestock, unintentional introduction of new species
Coexistence
Species can coexist in the same location if they don’t compete for the same resource
Fundamental Niche
Full range of condition or resources that a species can survive in (doesn’t take into account other species in the area)
Realized Niche
The actual set of conditions that the species live in the presence of other species (ex. birds separating into different level of the same tree due to competition)
Niche partitioning
Separation of niches: more niches partitioning, less competition, greater coexistence
Spatial Refuges
Simple environment: no refuges for prey, predator kills prey, then goes extinct
Complex environment: refuges for prey, prey is killed, but can survive in other places enough to escape predator
Disturbance
Any change in abiotic or biotic conditions (change in weather, new species, extinction)
Primary Succession
Community is empty, needs immigration to start again (ex. glacier retreat)
Secondary Succession
Populations decline but does not die out. Early-successional species are outcompeted by late-successional species
Scales of Diversity
Spacial Grain: characteristic scale (ex. 1×1 meter rectangle) Increase grain size=reducing pixel size
Spacial Extent: the overall region (an entire state) Increase extent=increase area of interest
Abundance
Number of individuals
Richness
Total number of species
Evenness
Relative equal distribution of all species
Composition
Identities of which species are present
LDG
Latitudinal Diversity Gradient
highest species richness near equator, lowest at the poles because more land area, less stress, more sun, higher temp, more time since no ice
Factors that increase richness over time
longer evolutionary time, larger area (island biogeography theory-larger and closer to mainland islands are better), longer time since disturbance, more indigenous land use, less poverty
Factors that decrease richness over time
More agricultural intensification (pesticide use), more land clearance (oil palms)
Agroforestry Systems
Maintain higher biodiversity than plantation agriculture, keeps natural landscape fragments, can also present TEK and cultural practices
Species Distribution
Affected by dispersal, abiotic/biotic, human activity, and behaviors
Dispersal
Movement of individuals and gametes to and from their original location
Often limited by behavior (birds refuse to cross gap due to fear of predators)
Abiotic and Biotic conditions on distribution
Abiotic: temperature and heat tolerance
Biotic: herbivory by cattle or competition between species
Environmental Gradients
Temperature, elevation, hurricane, predation risk gradients (could be continuous or patchy)
Biomes
Region experiencing similar environmental conditions and support a similar set of core species (Mojave and Sonoran Deserts), depends on climate, precipitation
Temperature Differences
Due to less sun at higher latitudes and more at lower latitudes, elevation causes air to expand and cool, less seasonality (cooler summer, warmer winter) near oceans
Precipitation Differences
Hadley cells leads to the tropics (mid-latitudes) to receive more rain, windward side of mountains facing ocean gets more rain than rain shadow on leeward side
NPP
NPP=GPP-R
NPP=net primary production
GPP=gross primary production
R=respiration rate
Climate drivers: water availability, extreme temperatures
Ecological Efficiency
Fraction of energy available to other organisms, average is 10%, causes trophic pyramids
Assimilation Fraction
Energy used by an organism for its own growth and respiration
Trophic Cascade
Bottom-up: limited resources determine producers and in turn upper trophic levels
Top-down: top predators determine energy flow and in turn lower levels
Metabolic rate of average person vs average plant
120 W, 4 W
Sociometabolism
Metabolism of humans accounting for bodily energy use and also indirect consumption such as agriculture, raising animals, burning biomass, fossil fuels, etc
Stock/Pool
Amount in the system
Flux
Rate of movement between compartments
Residence time
How long something spends in a equilibrium system (input=output), =stock/flux
Sink and Sources
Sink: positive net flux, increases
Source: negative net flux, decreases
Carbon Cycle
Atmosphere is not at equilibrium: net flux is positive, means greenhouse gases increase
Humans: fossil fuels burning (bigger impact), land use change (deforestation)
Nitrogen Cycle
Nitrogen builds DNA, RNA, ATP, etc.
Nitrogen reaches animals and plants through nitrogen fixing bacteria or decomposers (nitrogen gas into ammonium, nitrates), nitrogen escapes animals and plants through decomposers and denitrifying microbes
Humans increase nitrogen inputs via nitrogen fertilizers and acid rain
Hager-Bosch Process
Production of nitrate from nitrogen gas using fossil fuels, humans contribute 51% to nitrogen flux
Alternatives to fertilizer
Nitrogen fixing crops used as polyculture, or crop rotation (beans and peanuts)
Phosphorus Cycle
Phosphorus comes from rock weathering and phosphate rock formation, humans also input large amounts for fertilizer
Greenhouse Gases
Absorb IR radiation and re-emit it, trapping heat. CO2, N2O, CH4, O3, H2O
Positive Feedback Loop
More and more: ice feedback(warming, more ice melts, lower sunlight reflected albedo, more warming), vegetation feedback (warming, more trees die, less CO2 absorbed, more warming), cloud feedback 1(warming, more high altitude clouds, more IR absorbed, more warming)
Negative Feedback Loop
More leads to less: radiation feedback (warming, more IR radiation emitted, more cooling), cloud feedback 2 (warming, more tropical altitude clouds, more sunlight reflected to space, more cooling)
Deforestation causes drought
Less plants, more runoff, less aquifers, drier conditions downwind
RCP
Representative Concentration Pathway: scenarios that make predictions such as global warming and CO2 levels based on different trajectories of population growth, economic development, and carbon efficiency
Climate Change
Hot places get hotter, polar will get hotter, dry places get drier, wet places get wetter
Novel climates can push species beyond fundamental niche (range shifts)
Effects on humans: More extreme weather events, changing crop yields
Phenology
Changes in timing of seasonal events relating to organisms, often due to climate change
Disease
genetic, immune, diet, developmental, physical, environmental/chemical, infectious
Biological pest control
Use wasps to kill caterpillars on cotton instead of insecticide
Gene drive
Introduce infertility mutation into mosquitoes to cause extinction
Disease ecology used in biological warfare
Rinderpest (disease of cattle), Smallpox (disease of humans), cover smut (fungal disease of wheat), “Agent Orange” (mixture of herbicides)
Organisms can cause diseases
Algal blooms, secrete toxic compounds into water
Parasites, feeds and lives on a host and causes harm
Pathogen, a organism or virus that causes disease but not living in host
Transmission of Pathogens
Contact (direct movement from one host to another), vehicle transmission (indirect via water, dust, aerosols), vector transmission (indirect via another species)
Janzen/Connell Effect
Diseases/enemies that increase when their hosts have high-density populations cause negative density dependence (prevents host species from becoming very large in population)
Metapopulation
Patches each with populations that are linked by immigration/emigration. Diseases spread as metapopulations, and can re-emerge from other patches even if it’s gone from other patches.
Reservoirs for pathogens
Environmental: location where pathogens live when they are not parasitizing
Biotic reservoir: location where pathogens live when they are parasitizing
Zoonotic disease: diseases that normally don’t affect humans but could be transmitted (Lyme disease)
SIR Model
Susceptible → infectious → recovered (or dead)
Epidemic occurs if infectious individuals starts to increase (dl/dt>0)
dI/dt = beta*S*I - m*I
beta=transmission rate m=recovery rate S=number susceptible I=number infected
(S*beta)/m = R0
R0=1: epidemic stays the same size R0>1: epidemic grows R0<1:epidemic shrinks
Many exceptions are more complicated than SIR model, like bubonic plague
Global changes that increase disease spread
Habitat destruction, urbanization, air travel, climate change
Indigenous fire management
Regular burning to promote habitat for food plants, game animals, etc. US uses fire suppression instead, that causes greater forest densities and larger fires
Pyromes
Fire-dependent biomes: Savanna, Chaparral, Coniferous forests
Plant adaptations to fire
Above-ground survival (thick bark, tall trunks), below-ground survival (resprouting), generation from seed (seed dormancy, post-fire germination), flammability, serotonin (cones or seeds survive fire and open after fire)
Anthromes
Human-modified landscapes (>70% of land)
Biotic Homogenization
Increases in similarity of communities over time (caused by increased dispersal of common species, increased mortality of rare species, human selection of species, and urbanization)
Dispersal Corridors
Protected areas are connected by them to promote persistence of species metapopulation across patches
Translocation/assisted migration
Humans move genotypes of species to new locations, but its very time/money-intensive
Habitat Restoration
Humans manipulate abiotic/biotic conditions and promote desirable succession pathways
Values of biodiversity
Intrinsic Value: existence value (the species exists), intrinsic (other species have a right to exist)
Instrumental value: reaches human goals/desires (non-market consumption, exchange, amenity, scientific/educational)
Cultural value: bald eagle is a symbol for nation, bison as a part of native american cultures
Service value: shade, food impact reduction
Ecosystem services
Provisioning: products generated by nature
Regulating: improving life by cleaning air and preventing erosion
Cultural: contribution to knowledge and society
Supporting: maintenance of basic life processes
Protected Areas
Bigger area, more species
Economics on deciding nature protection
Economic frames nature by assigning money-based values instead of natural capital
Pros: gives common language for decision-making, could be more effective than community or government protests due to strength of market, comfortable framing for corporation and governments
Cons: assumes capitalism is all that matters, assumes buying/selling nature is ethical, neglects long-term effects but rather short-term changes in market, enables powerful and rich ppl to “pay to pollute”
The right framing of biodiversity
If humans have natural rights, then nature also has rights, not just a resource only used by humans, humans have responsibility to care for nature