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What is marine?
Having to do with the ocean
What is ecology?
The study of the relationships of organisms with each other and with their physical environment
What is conservation?
The act of protecting Earth's natural resources for current and future generations
How do the spatial dimensions of life on land compare to those in the ocean?
Terrestrial: thin veneer of life along surface
Ocean: life distributed across large depth range
Surface area: Oceans cover ~70% of Earth, land ~30%.
Depth of life:
Land: life is mostly limited to soils, plants, and low-flying organisms → ~36 m average.
Ocean: average depth of ~3,600 m where life is found.
Comparison: The ocean is ~100x deeper for life than land.
Overall space for life: When depth is combined with surface area, the ocean provides ~200 times more space for life than land.
How is light distributed across Earth, and why does it vary?
Light is unevenly distributed and shifts with the seasons due to Earth’s tilt.
Poles: strongest seasonal effect → up to 24 hours of daylight in summer and 0 hours in winter.
Equator: light is consistent → ~12 hours of daylight year-round.
Angle of sunlight: most direct at the equator → light is more intense compared to higher latitudes.
What happens to sunlight when it enters the ocean?
Sunlight is made of many colors (visible in a prism).
Once it hits the ocean, light is reflected or absorbed.
Water acts as a filter:
Red light is absorbed first (shallowest).
Blue light penetrates the deepest.
If you were to dive deep into the ocean, what colors would you see?
Blue, bc it travels farthest
What type of radiation is visible light?
electromagnetic radiation, classified by wavelength.
Are most types of light visible to humans?
No. Most electromagnetic radiation is not visible to us, such as:
Ultraviolet (UV)
Infrared (IR)
Why is visible light important in the ocean?
best wavelengths for penetrating water.
In contrast, UV and IR do not penetrate water well—water is essentially opaque to them (like black coffee is to visible light).
Explain how Depth and Light work in the ocean.
Because light gradually gets absorbed or scattered by ocean water, it gets darker as you travel deeper
What are the ocean light zones by depth?
Photic zone (with sunlight)
Euphotic: enough light for photosynthesis
Dysphotic: dim light, not enough for photosynthesis
Aphotic zone: no sunlight
What is important to know about the relationship between ligh in the ocean?
Some colors of light penetrate deeper into the ocean
How might the depth distribution of light in the ocean affect life?
Photosynthesis near surface
• Life concentrated in shallows
• Color of organisms can change with depth
• Ecosystems in dim light or dark may depend on food from above
• Bioluminescence in the deep ocean
Why is the Ocean blue?
Some light that
enters the ocean is
scattered and
bounces back to the
surface, creating its
color
• Most of the light that
bounces back is blue,
because the other
colors have been
absorbed
Water Motion: What is the difference between standing waist deep in a 10mph stream vs. standing in a field with 10 mph wind on a sunny day?
Force pushing you
• Related to density of water
(water has a greater
density)
• Same speed has a very
different effect
How might water motion and density affect biology?
• Moving life that drifts in the
flow
• Life concentrates where
nutrients and food are
delivered
• Some organisms anchored to
the bottom
• Some organisms have evolved
of hydrodynamic shapes
• Water flow affects local
temperature
What are the primary causes of water motion?
Wind
• Tides
• Density differences
• Earth’s rotation
What are ocean gyres and why are they important?
large circular surface currents in the ocean
Driven by global wind patterns + Earth’s rotation (Coriolis effect)
Found in each major ocean basin
Important for:
Transporting heat, nutrients, and organisms
Concentrating debris (e.g., Great Pacific Garbage Patch)
Upwelling
the delivery of deep water to
the surface, which is often
• Colder
• Nutrient rich
How do Tides effect the motion of water?
Gravitational pull of
moon and sun
• Larger tides when
they’re lined up
• Smaller tides when
out of alignment
How might tides affect life?
Exposure to air at low tide
Drive some coastal currents
How are wave formed?
by surface winds
Friction between wind and ocean surface creates ripples, which grow to waves
What are wind waves?
small, irregular waves that are made locally by wind
What does Swell mean.
Is waves that started as wind
waves, but travel beyond the
source of the wind
• Usually regular and smooth
• Can occur when it’s not windy
• Can travel across ocean basins
Why are Waves important for ecologically?
• Mix nutrients and other materials vertically
• Move materials laterally near shore, as they break
• Structure ecosystems by differentially removing fragile organisms
When do waves break?
When they reach shallow water
Which "waves” are not cause by wind?
Tsunamis: which are caused by land motion underwater – bathtub effect
Tidal waves: formed by tidal currents
What might deeper water be colder?
Liquid can become stratified, with stable layers with different densities and other properties
• Examples
• Thermocline: warm water
floats on cold water
• Halocline: Less salty water
floats on saltier water
What does Stratification do?
Can trap some water well below the surface, with upper layers capping those below
Organic matter can sink to deep water where it decomposes, trapping nutrients
What kind of waves mix deeper?
Big waves
What does the depth of mixing depend partly on?
Stratification
Weak stratification allows deeper
mixing
How does latitude affect ocean mixing, and why is this important for nutrients?
Warm regions: strong thermocline → mixing is relatively shallow
Cold regions: weaker thermocline → mixing is deeper
Importance: deeper mixing brings nutrients to the surface, supporting primary production
What drives Global Surface Currents?
Coriolis force drives currents
• Clockwise in north
• Counterclockwise in south
Create circular patterns called
gyres
Details change a smaller scales
due to factors such as
• Bottom topography
• Land
• Other currents
• Local winds
What is driven by the Coriolis effect?
Ekman Transport
Northern hemisphere example:
• Water moves with wind a surface
• Friction between water layers pulls
deeper water along
• Water curves to the right with depth
• Net water movement 90 degrees to
right of wind
Explain how Coastal upwelling happens?
• Along-shore winds create
surface currents
• If going the right direction,
along-shore surface currents
transport water offshore
• What replaces the water
that moves offshore?
• Hole/negative space near
shore
• Deep water is pulled to
the surface to fill the hole
Coastal Upwelling Zones
Concentrated on
west coasts
- Equator-ward
winds cause
offshore transport
- Cool, nutrient rich
upwelled water
Results in:
• Cool air / fog
• High nutrient
concentrations
What are other causes of upwelling?
• Currents interacting with
topography, for example,
hitting oceanic islands
• Surface mixing from
waves and tides
• Equatorial upwelling
• Similar mechanism to
coastal upwelling
• But happens in two
directions with equator
acting like ‘shore’
• Thermohaline circulation
What is Thermohaline Circulation?
Driven by changes in
density
• Temperature
(thermo)
• Salinity (haline)
As salty water cools in
the North Atlantic, it
sinks
Pushes deep water out
of its way, driving a slow
global current
How does the Thermohaline Circulation work?
Gulf stream brings warm surface
water to North Atlantic
• Water cools in cold wind/air
• Eventually, ice forms
• Brine rejection: ice formation rejects
salt, leaving remaining water saltier
Surface water is more dense: colder
and saltier
• It sinks
• Pushes other deep water, driving a
slow global current
Why is the Thermohaline Circulation considered the World’s largest current?
Connects all major ocean basins
Deep water accumulates nutrients as surface material sinks
Redistributes nutrients and heat globally
What’s the most important chemical reaction in biology?
My opinion: photosynthesis
Simplified photosynthesis equation:
• Energy (sunlight) + CO2 + H2O ↔ Carbohydrate + O2
What does this equation mean: • Energy (sunlight) + CO2 + H2O ↔ Carbohydrate + O2
Equation goes both ways:
• Going right is photosynthesis:
creation of carbohydrates
• Going left is respiration:
breakdown of carbohydrates
What is a Primary Production (Autotrophy)?
creation of organic matter from inorganic carbon (CO₂) using energy
Autotrophs = organisms that produce their own food
Examples: phytoplankton, algae, seagrasses
Importance:
Base of the marine food web
Supports all higher trophic levels
Drives carbon cycling in the ocean
What are Primary Producers (Autotrophs)?
are organisms that can do things like Photosynthesis.
Are there any other forms of Primary Production besides photosynthesis?
• Chemosynthesis
• Main differences is that the energy source is chemical rather than light
• Examples include hydrogen gas or hydrogen sulfide
Chemosynthesis in Hydrothermal Vents
Hydrothermal vents are deep
ocean fissures where superheated
water, with chemicals like
hydrogen sulfide, enters the ocean
Some kinds of microbes can use
inorganic chemicals (e.g.,
hydrogen sulfide) in the hot water
to fuel primary production
without sunlight
A diverse ecosystem can be
formed based on chemosynthesis
Explain what Eukaryotes are?
• Have cells with a nucleus
• Familiar plants, animals, fungi
• Protists: everything else
• Grab bag of different groups including algae and protozoa
What are Prokaryotes?
No nucleus
• DNA is circular
• All single cellular
• Smaller than eukaryotes
What is Bacteria?
Some autotrophic
What characteristics make a Archaea?
Many live in extreme environments
• Some autotrophic
• Evolutionarily closer to us than
bacteria!
What are the main groups of Primary Producers in the Ocean?
Plants
• Algae
• Prokaryotes (Archaea and
Bacteria)
Plants in the Ocean?
Plants have vascular tissues like roots and stems
In the ocean, plants are confined to shallow coastal areas
Main groups:
• Mangrove trees:
• Leaves and stems above water
• Roots underwater at high tide
• Seagrasses:
• Resemble grasses on land
• Live in shallow coastal areas
• Can store large amounts of carbon in sediments
• Salt marsh plants, live in estuaries (places where fresh and
saltwater mix)
Plants are locally important in the ocean, but minor players globally
What are Algae?
Relatives of plants that lack roots and
stems
Abundant and diverse in the ocean
What are the two main functional groups that Algae have?
Macroalgae (AKA seaweed)
Multicellular, plant-like algae
Includes kelps
Mainly anchor to the substrate
Plants evolved from green macroalgae
Microalgae
Small, often unicellular, primary producers
Can grow in films on the bottom
Most float in water
What are Prokaryotes?
• Microscopic single celled organisms
• Archaea and bacteria
• Some are primary producers
What are Phytoplankton?
• The small primary producers that are
adrift in water
• Includes both microalgae and
prokaryotes
• Responsible for most of the primary
production in the ocean
Why are most primary producers in the ocean microscopic, unlike land, which is dominated by larger plants?
Possible answers:
• Surface area to volume ratio: easier to absorb
nutrients from environment when small
• Rapid response to changing conditions: when
nutrients or sunlight arrive, microscopic
organisms can quickly grow their populations
• Bet hedging: by being small and abundant,
chances are better that some of your relatives
will find good conditions
• Little advantage to being big:
• Can’t anchor to hold space for competition
• No need for roots to gather nutrients
• Neutral buoyancy: no need for hard structures to
support growth
What are the important nutrient needed for Photosynthetic Primary Production?
• Nitrogen
• Phosphorous
• Iron
When all needs of primary producers are
met, growth can be extremely fast
What limits primary production?
• Water: Always present
• CO2: Rarely limiting in ocean
• Light:
• Large influence by depth
• Influenced by latitude,
strong seasonal pattern
• Nutrients: frequently the
limiting factor in surface
water
• Used up quickly
• Sink beyond light
• Source needed for high
productivity
What is the purpose of a Trophic Status
To classify Aquatic ecosystems on the basis of how much primary productivity they support
What is the Trophic Status of the level: Oligotrophic?
low nutrient levels and low primary production
What is the Trophic Status of the level: Mesotrophic?
medium nutrient levels and primary production
What is the Trophic Status of the level: Eutrophic
high nutrient levels and high primary production
Give an example of how Primary Production fuels ecosystems?
Arctic example
• Summer
• High light
• High nutrients
• Respiration over winter
• Strong mixing once ice is gone
• Phytoplankton blooms
• Small animals eat phytoplankton
• Fuels growth of larger animals
Bottom line: whales, polar bears and
walruses rely on the nutrients that fuel
phytoplankton growth
This is Secondary Production
Define Heterotrophs?
are organisms that use energy stored by autotrophs to fuel life
Define Secondary production?
the process by which heterotrophs convert organic molecules from primary producers into new life

Describe the Classic Ocean Foodweb?
Built from the bottom up:
• Primary producers supply food
• Eaten by small grazers
• Eaten by larger predators
• Etc.
• Key concepts
• Herbivore/grazer: eats only
autotrophs
• Carnivore: eats only animals
• Omnivore: eats primary
producers and animals

What are the different Trophic Levels?
1. Primary Producers
2. Herbivores
3. Intermediate predators /
carnivores
4. Top predators / carnivores
Every time an organism eats another, energy is lost. Why?
• Heat
• Metabolism
• Incomplete digestion
• Reproduction
What is Conversion efficiency?
a measure of how much biomass (the weight of life) is transferred from one trophic level to the next
What’s the trophic level of a carnivore that eats only herbivores?
Level 3: Intermediate predators / carnivores
What’s the tropic level of autotrophs?
Level 1: Primary Producers
What’s the tropic level of an omnivore that eats both autotrophs and herbivores?
Between 2 and 3
What are the Major groups of heterotrophs?
Entirely heterotrophic groups
• Animals
• Fungi
Some species are heterotrophic
• Protists
• Bacteria
• Archaea
What are Microscopic Heterotrophs in the Ocean?
Protists
• All eukaryotes except plants,
animals, slime molds or fungi
• Diverse grab-bag of species
• Includes autotrophic algae, single
celled amoeba
Bacteria
• Single celled prokaryotes
Archaea
• Single celled prokaryotes
• Evolutionarily distant from
bacteria
Note: some protists, bacteria, and
archaea can be both heterotrophic
and autotropic at the same time
What do microscopic heterotrophs eat?
DOM: Dissolved Organic Matter
• POM: Particulate Organic Matter
Bacteria + Archaea
• So small that they cannot ingest food (too
small to have a mouth)
• Osmotrophs: organisms that consume
dissolved food
• Can also use enzymes to dissolve POM for
digestion
Microzooplankton, like some protist
• Consume POM through ingestion
Micro-organisms in the Ocean?
Standard ocean food chain:
small organism consumed by
progressively large ones
But much of the primary
production doesn’t make its
way directly up the food chain
due to imperfect conversion
efficiency
But, it’s not necessarily lost…
Describe the The Microbial Loop?
Many primary producers die and
leak before being eaten
Some of their biological molecules
dissolve into the water
Heterotrophic archaea and
bacteria, consume these
molecules and grow
Micro zooplankton, comprised of
tiny animals and protists, eat the
archaea and bacteria
Larger animals eat the smaller
ones
This is called the microbial loop
Archaea and bacteria also release
nutrients back into the water,
fueling more primary production
What are the microbes in the microbial loop?
• Defined unicellular animals that
are too small to see without a
microscope
• Includes
• All bacteria and archaea
• Single cellular protists and
fungi
• In this diagram can be
• Primary producers
• Heterotrophic archaea and
bacteria
• Microzooplankton
What is the importance of Micro-organisms in the Ocean?
The microbial loop increases the conversion efficiency of marine food webs: more energy makes it to higher trophic levels Nutrient recycling boosts primary productivity None of this is visible to the eye
What would happen to ocean ecosystems without osmotrophic microbes?
Osmotrophic microbes (bacteria and archaea) break down dissolved organic matter and recycle nutrients.
Without them:
Organic matter would accumulate, leading to nutrient depletion for other organisms.
Primary production would decline due to lack of recycled nutrients.
Overall food web efficiency would drop, impacting all levels from plankton to fish.
Essentially, biogeochemical cycling in the ocean would collapse, destabilizing ecosystems.
Review of Ocean Processes
Physics sets the stage for biology,
driving properties like
• Light availability
• Water motion
• Nutrient distribution
• Water chemistry
Life responds to these
environmental conditions
• All life starts with primary
production
• Secondary consumers feed off
primary producers and other
heterotrophs to build
foodwebs
Whare are the four scales of organization in ecology discussed in lecture?
• Individual: one organism
• Population: a group of
interacting organisms of
the same species
• Community: a group of species that are actively
interacting with each
other at the same time
and place
• Ecosystem: a community
of organisms and the
surrounding physical
environment
Define the scale Individual?
one organism
Define Population?
a group of interacting organisms of the same species
Define Community?
a group of species that are actively interacting with each other at the same time and place
Define Ecosystem?
a community of organisms and the surrounding physical environment
What is a niche?
Organisms are adapted to live in certain environments
the set of environmental and biological conditions in which an organism can live
What is an example of a niche?
Crabeater seal example
• Lives only around Antarctica
• Requires access to places to
haul out for resting and giving
birth
• Eats only krill
• Harbor seal example
• Lives northern Pacific and
Atlantic
• Requires access to places to
haul out for resting and giving
birth
• Eats fish, mollusks,
crustacean
In terms of diet, which one has the relatively broader niche? The harbor seal or the crabeater seal?
• Generalists: relatively broad niche (harbor seal)
• Specialists: relatively narrow niche (crabeater seal)
What are some implications of having large changes in body size during development?
Many ocean organisms undergo extraordinary shifts in body
size
• This means that they must shift food with growth
• Goliath grouper example:
• Adult eats fish, adult lobsters, even sea turtles!
• Juvenile: eats tiny plankton
What is a developmental phase that many marine organisms have?
planktonic larvae
What does Plankton mean?
generally small aquatic organisms that are at the mercy of ocean currents
What does Larvae mean?
early developmental stages of organisms, not yet able to reproduce
why do you think so many marine organisms have planktonic larvae?
• If you live near structure
(reef, coastline, etc.),
there are often many
predators
• Small offspring are not
safe
• It’s better to get them
into open water where
predators are often rarer
• Open water also tends to
have small food items for
small animals
How do you as a larvae avoid being eaten in the open ocean?
• One strategy:
• Be small and clear in a big space
• Like a contact lens in a swimming pool
can you think of any ecological consequences of having planktonic larvae?
• Examples:
• Offspring drift away
• May not live near parents
• Can colonize new places
• High death rate for planktonic larvae that don’t drift to the
right places
• May have high variability in success of offspring
• Good years / bad years due to chance
Define Metamorphosis
Major and rapid transition in bodily form and function

How do Marine organisms perform metamorphosis?
• Settlement: The transition from
plankton to benthic (bottom) habitat
• Settlement involves metamorphosis
• Example: flatfish: sole, flounder,
halibut