Acoelomate
Without a coelom, as in flat worms and proboscis worms
Adaptive radiation
Evolutionary diversification that produces numerous ecologically disparate lineages from a single ancestral one, especially when this diversification occurs within a short interval of geologic time
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Zoology
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Acoelomate
Without a coelom, as in flat worms and proboscis worms
Adaptive radiation
Evolutionary diversification that produces numerous ecologically disparate lineages from a single ancestral one, especially when this diversification occurs within a short interval of geologic time
Allele
Alternative forms of genes coding for the same trait, and situated at the same locus in homologous chromosomes
Allelic frequency
An estimation of the proportion of gametes produced in a population (gene pool) that contains a particular allelic form of a particular gene
Blastocoel
The cavity within the hollow mass of embryological cells
Blastomere
An early cleavage cell
Blastopore
External opening of the archenteron in the gastrula.
Blastula
Early embryological state of many animals; consists of a hollow mass of cells
Carnivore
Any organism that eats animals
Carrying capacity
The maximum number of individuals of a particular species that can persist under specified environmental conditions
Cephalization
The evolutionary process by which specialization, particularly of the sensory organs and appendages, became localized in the head end of animals
Clade
A taxon or other group consisting of a particular ancestral lineage and all of its descendants, forming a distinct branch on the cladogram or phylogenetic tree
Cladistics
A system of arranging taxa by analysis of primitive and derived characteristics so that the arrangement reflects phylogenetic relationships
Cladogram
A branching diagram showing the pattern of sharing of evolutionarily derived characters among species
Coelom
The body cavity in triploblastic animals, lined with mesodermal peritoneum
Commensalism
A relationship in which one individual lives close to or on another and benefits, and the host is unaffected; often symbiotic
Common Descent
Darwin’s theory that all forms of life are derived from a shared ancestral population through a branching of evolutionary lineages
Competition
Some degree of overlap in ecological niches of two populations in the same community, such that both depend on the same food source, shelter, or other resources, and negatively affect each other’s survival
Competitive exclusion
An ecological principle stating that two species whose niches are very similar cannot coexist indefinitely in the same community; one species is driven to extinction by competition between them
Decomposer
A consumer that breaks organic matter into soluble components available to plants at the base of the food web; most are bacteria or fungi
Definitive host
The host in which sexual reproduction of a symbiont occurs; if no sexual reproduction, then the host in which the symbiont becomes mature and reproduces
Deme
A local population of closely related animals
Demography
The properties of the rate of growth and the age structure of populations
Density-dependent
Biotic environmental factors, such as predators and parasites, whose effects on a population vary according to the number of organisms in the population
Density-independent
Abiotic environmental factors, such as fires, floods, and temperature changes, whose effects on a population are unaffected by the number of organisms in the population
Derived character state
Condition of a taxonomic character inferred by cladistics analysis to have arisen within a taxon being examined cladistically rather than having been inherited from the most recent common ancestor of all members of the taxon
Deuterostomia
A group of higher phyla in which cleavage is indeterminate and ancestrally radial. The endomesoderm is enterocoelous, and the mouth is derived away from the blastopore
Dioecious
Having male and female organs in separate individuals
Ectoderm
Outer layer of cells of an early embryo (gastrula stage); one of the germ layers, also sometimes used to include tissues derived from ectoderm
Ectoparasite
Parasite that resides on the outside surface of its host organism
Endoderm
Innermost germ layer of an embryo, forming the primitive gut
Endoparasite
Parasite that resides inside the body of its host organism
Evolution
All changes in the characteristics and diversity of life on earth throughout history
Evolutionary species concept
A single lineage of ancestral-descendant populations that maintains its identity from other such lineages and has its own evolutionary tendencies and historical fate
Evolutionary taxonomy
System of classification, formalized by George Gaylord Simpson, that groups species into Linnean higher taxa representing a hierarchy of distinct adaptive zones; such taxa may be monophyletic or paraphyletic but not polyphyletic
Frontal plane
Plane parallel to the main axis of the body and at right angles to the sagittal plane
Fundamental niche
A variety of roles potentially performed by an organism or population in an ecological community; limits on such roles are set by the intrinsic biological attributes of an organism or population
Gastrocoel
Embryonic cavity forming in gastrulation that becomes the adult gut; also called an archenteron
Gastrula
Embryonic stage, usually cap- or sac-shaped, with walls of two layers of cells surrounding a cavity with one opening (blastopore)
Gene pool
Collection of all the alleles of all the genes in a population
Gradualism
A component of Darwin’s evolutionary theory postulating that evolution occurs by the temporal accumulation of small, incremental changes by populations, usually across very long periods of geological time
Guild
Species of a local community that partition recourses through character displacement to avoid niche overlap and competition, such as Galapagos finch communities whose component species differ in beak size for specializing on different-sized seeds
Herbivore
Any organism subsisting on plants
Homeothermic
Having a nearly uniform body temperature, regulated independent of the environmental temperature
Homology
Similarity of parts or organs of different organisms caused by evolutionary derivation from a corresponding part or organ in a common ancestor, and usually having a similar embryonic origin
Hypothesis
A statement or proposition that can be tested by observation or experiment
Inheritance of acquired characteristics
The discredited Lamarckian notion that organisms, by striving to meet the demands of their environments, obtain new adaptions and pass them by heredity to their offspring
Intrinsic growth rate
Exponential growth rate of a population; the difference between the density-independent components of the birth and death rates of a natural population with stable age distribution
Limiting resource
A particular source of nutrition, energy, or living space whose scarcity is causally associated with a population having fewer individuals than otherwise expected in a particular environment
Macroevolution
Evolutionary change on a grand scale, encompassing the origin of novel designs, evolutionary trends, adaptive radiation, and mass extinction
Mesoderm
The third germ layer, formed in the gastrula between the ectoderm and endoderm; gives rise to connective tissues, muscle, urogenital and vascular systems, and the peritoneum
Metapopulation dynamics
The structure of a large population that comprises numerous semi-autonomous subpopulations, termed demes, with some limited movement of individuals among demes. Demes are often geographically distinct
Microevolution
A change in the gene pool of a population across generations
Monoecious
Having both male and female gonads in the same organism, usually denoting that this is the typical condition of a species; hermaphroditic
Monophyly
Condition occurring when a taxon or other group of organisms contains the most recent common ancestor of the group and all of its descendants
Ontogeny
The course of development of an individual from egg to senescence
Paraphyly
The condition that a taxon or other group of organisms contains the most recent common ancestor of all members of the group but excludes some descendants of that ancestor
Parasitism
The condition of an organism living in or on another organism (host) at whose expense the organism is maintained
Parasitoid
Organism that kills the host during or at the completion of development
Parenchyma
In simpler animals, a spongy mass of vacuolated mesenchyme cells filling spaces between viscera, muscles, or epithelia
Phenotype
The visible or expressed characteristics of an organism: influenced by the genotype in interaction with environmental conditions
Phyletic gradualism
A model of evolution in which morphological evolutionary change is continuous and incremental and occurs mainly within unbranched species or lineages over long periods of geological time
Phylogenetic tree
A diagram whose branches represent evolutionary lineages; depicts the common descent of a species or higher taxa
Phylogeny
The origin and diversification of any taxon, or the evolutionary history of its origin and diversification, usually presented as a dendrogram
Polyphyletic
The condition that a taxon or other group of organisms is derived from more than one ancestral source
Pseudocoelom
Body cavity not lined with peritoneum and not a part of the blood or digestive systems; embryonically derived from the blastocoel
Punctuated equilibrium
Model of evolution in which morphological evolutionary change is discontinuous, being associated primarily with discrete, geologically instantaneous events of speciation that lead to phylogenetic branching; morphological evolutionary stasis characterizes species between episodes of speciation
Radial cleavage
Embryonic development in which early cleavage planes are symmetrical to the polar axis, each blastomere of one tier lying directly above the corresponding blastomere of the next layer
Realized niche
The role actually performed by an organism or population in its ecological community at a particular time and place as constrained by both its intrinsic biological attributes and its particular environmental conditions
Reproductive barrier
The factors that prevent a sexually propagating population from interbreeding and exchanging genes with another population
Resource
Available source of nutrition, energy, or space in which to live
Resource partitioning
To coexist in the same habitat, two or more species specialize on different portions of a shared resource
Sagittal plane
Pertaining to the median anteroposterior plane that divides a bilaterally symmetrical organism into right and left halves
Sink deme
A subpopulation whose members are drawn disproportionately from other subpopulations of the same species because of unstable environmental conditions resulting in the destruction of individuals in the subpopulation and then subpopulation is replenished by colonists from other subpopulations when favorable conditions are restored
Source deme
A stable subpopulation that serves differentially as a source of colonists for establishing, joining, or replacing other such subpopulations of the same species in nearby environmentally unstable areas
Species diversity
The number of different species that coexist at a given time and place to form an ecological community
Species richness
The number of different species that coexist at a given time and place to form an ecological community
Spiral cleavage
Type of early embryonic cleavage in which cleavage planes are diagonal to the polar axis, and unequal cells are produced by the alternate clockwise and counterclockwise cleavage around the axis of polarity
Survivorship
The proportion of individuals of a cohort or population that persist from one point in their life history, such as birth, to another one, such as reproductive maturity or a specified age
Systematics
Science of taxonomy and reconstruction of phylogeny
Systematization
Construction of a hierarchical taxonomy that conveys the structure of common evolutionary descent among species. Each recognized taxon comprises an ancestral population lineage and all of its descendants
Transverse plane
A plane or section that lies or passes across a body or structure dividing it into cephalic and caudal pieces
Variation
Differences among individuals of a group or species that cannot be ascribed to age, sex, or position in the life cycle
Velarium
Shelf-like extension of the subumbrella edge in cubozoans
Velum
A membrane on the subumbrella surface of jellyfishes of class Hydrozoa