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Behavior
the action or reaction of individuals in response to a stimulus (foraging, repro, maintainace, pred avoid, exploration)
proximate causes (factors)
machanisms within the individual that operate to make the behavior possible (genetic, development, physioloigical)
ultimate factors
evolutionary causes (why has the animal evoloved those mechanisms, selective advantage)
ethology
study of animal behavior
behavioral ecology
study of the inter-relationships between animal behavior and environment
sociobiology
study of social behavior in animals; evolution, genetics, population biology, ecology
innate behavior
developmentally fixed, instinctive, not based on prior experience (FAPs = fixed aciton pattern)
learning
experienced-based modification of behavior
Maturation
changes in behavior caused by developmental changes in neuromuscular systems (flight)
associative learning
associating one stimulus with another
classical conditioning
associating arbritrary stimulus with reward or punishment
operant conditioning
trial and error learning (training)
importance for senses in vertebrates
being able to detect variability of the environment (define and limit known world)
function of sense organs
monitor the environment, permit evolution of complex behaviors
chemoreceptors
detect smell and taste, most ancestral and universal, smell is more sensitive than taste
taste
taste receptor cells (TRC) located in taste buds, which are located mainly on tongue in mammals but also in other areas that contact food (five basic tastes: sour, salty, bitter, sweet, umami)
smell
more complex than taste, receptors located in nasal cavity (20,000,000 olfactory receptors in human nose)
important functions of chemical scents
recognition, emotional states, advertisement, orientation
mechanoreceptors
touch/pain, hearing/lateral line, equilibrium
receptors for touch and pain
receptors located in the skin, the receptor is distorted and then local flow of electrial current, then info relayed to CNS, most touch receptors are in the face and limbs, pain receptors may response directly or indirectlu to injury
receptors for pressure and sound
lateral line in fish and aquatic amphibains decects pressure from waves and vibrations in the water, important in schooling (sensory cells in pits and grooves, hair cells in cupula sense pressrue); hearing, sensing vibrations of frequencies (external→fluid→membrane→move sensory cells→nerve impluses to CNS)
balance
vertebrate ear originated as a _______ organ (two small chambers and 3 semicircular canals give info about head position and movement, detect changes in position)
mammalian hearing
ear w/ 3 divisions: outer, middle, and inner ear; outer ear collects sound waves to take to tympanic membrane, middle hear has air filled chamber with 3 bones that amplify the vibrations from tympanic membrane, inner ear has liquid-filled cochlea where hair cells become displaced with vibrations
pitch discrimination and loudness in mammalian hearing
located of hairs stimulated and number of hairs being stimulated
adaptations for hearing
enlarged middle ears, asymmetrical ears
electromagnetic reception
light, heat, electrical/magnetic signals
rods and cones
receptors for light (photoreceptors)
eyeball
3 layers (sclera, choroid, retina)
cornea
transparent, bends light in the eye to have it reach retina
Iris
regulates the opening for light (pupil)
lens
helps to focus the light
retina
contains photoreceptor cells (rods and cones)
rods
achromatic vision, lower visual acuity, good for vision in dim light, contain rhodopsin
cones
color discrimination, good for seeing in light conditions, contain photopigments to sense red, green, blue
diurnal
many ____ animals can see in color, nocturnal animals are color blind
350 MYA
color vision evolved in vertebrates around _________.
median eye
light sensitive spot on the top of the head in many fish and some reptiles (image forming eyes)
lateral eyes
have monocular vision, no overlap of vision fields of left and right eyes (large vision field but cannot distingush distance)
frontal eyes
binocular vision, large overlap of vision fields, very good depth perception, reduced field of view
adaptation for amphibious fish
two pupils/eye with aquatic and aerial vision
round pupils
only nocturnal or diurnal
vertical pupil
nocturnal and sometimes active during the day
vertical pinhole
Several images, yield great depth of field despite monocular vision
horizontal pupil
Allows wide-angle scanning. Good in wide-open habitats
Nictitating membrane
transparent membrane that helps to clean and prevent the eye drying
pit organs
an organ found on snakes to detect heat
electromagnetic receptors
fish can detect electric signals through a network of electrosensitive cells (fish can emit and recieve these signals)
Major functions of the nervous system
The controlling, regulatory, and communicating sytem in the body; in conjuntion with the endocrine system is to maintain body function and homeostasis (fast-acting)
Central Nervous System
brain and spinal cord
Peripheral nercous system
all peripheral nerves
Neuron
basic cellular unit of the nervous system
Dendrites
branched receptors of the neuron to recieve signals
Axon
long, and carries impluse away from the cell body
Sensory neurons (afferent)
sense the environment and take sensory info to the CNS
Interneurons (99% of all neurons)
Connect neurons together in CNS (intergrate info)
Motor neurons (efferent)
Recieve info from CNS and takes it to effector organs to trigger a response
Characteristics of a nerve impulse
occur in response to stimulus, produced by changes in membrane potential, all-or-none responses, signal intensity is signaled by frequency of impulses
Schwann Cell
Produce insulating mylin sheath to help maintain impulse down the axon
How an impulse is produced
resting potentials (high K+ inside axon and Na+ outside axon maintained by pump), voltage change opens Na+ permeable channels and membrane depolarizes, repolarizes when K+ channels open
Characteristics of nerve propogation
small axons conduct slowly, axons are bigger where fast responses are needed, vertebrates increase propagation speed with myelination (charge propagates by saltatory movement)
Nerve synapse
sites where signals are transmitted from one neuron to another (electrical and chemical)
Electrical synapse
charge flows across “gap-junction”, no time lag (therefore very fast), good for escape reactions
Chemical synapse
use specialized neurotransmitters that diffuse from one membrane to the other to propagate signal
Invertbrate nervous system
not complex, very basic (nerve nets rather than central nervous system)
Vertebrate nervous systems
Nerves interconnecting spinal cord and brain, spinal cord hasn’t changed much over evolution, cerebrum becomes elaborated further down the lineages
increase
Brains ________ with body size (but not a 1:1 ratio)
olfactory
Birds’ _______ region in the brain is greatly reduced
optic
Birds’ _______ lobes are enlarged
developed
In mammals the olfactory is very ________
_______ can generate fresh supplies of brain cells
Adult canaries and zebra zinches
Traits of the peripherial nervous system
Motor and sensory neurons are bundled, two subdivisions (somatic and autonomic)
Autonomic nervous system
Made up of Parasympathetic and sympathetic systems (maintain balance of homeostasis)
Parasympathetic system
lowers heart rate and respiritory rates
sympathetic system
increases heart rate and respiratory rate
Somatic nervous system
innervates skeletal muscles for movement impulses
Reflex arc
neurological feedback loops, only 2 or 3 neurons, bypasses brain
70-80% water
vertebrate bodies contain __________.
salt ions
Many ________ important for body function are dissolved in water.
Water balance
keeping from drying out or getting flooded
Salt balance
keeping from getting to salty or not salty enough
Osmosis
movement of water along solute concentrations
Kidneys
Key osmoregulator; main organ for excretion of nitrogenous wastes (filtration, reabsorbtion, secretion)
Nephron
Functional unit of the kidney; contains glomerulus (filter), Bowmans capsule (collects ultrafiltrate), proximal tubule (site of reabsorbtion), Loop of Henle (water reabsorbtion and release of excess ions), distal tubule (site of waste secretion), collecting ducts (collects urine and directs it to urinary system, controls the concentration)
glomerulus
filters the blood, leaky capillaries produce ultrafiltrate
Bowmans capsule
site of collection for ultrafiltrate
proximal tubule
site of reabsorbtion for nutrients and ions
Loop of Henle
Descending limb: main site of water resorbtion, ascending limb: active removal of exvess ions
Distal tubule
site of waste secretion (nitrogenous wastes, urine volume reduction)
Collecting ducts
collects urine, controls concentration, sends to urinary system
Discharged through urethra
where does the urine exit the body in mammals
Birds have _____ urinary bladders?
zero (adaptation for flight)
Solutions of marine fish to maintain osmotic balance
1) Be as salty as the environment (hagfish) 2) Have as high osmotic pressure as the environment (cartilagenous fish using urea) 3) Have mechanisms to counteract water loss and salt gain (bony fish using chloride cells to pump out salt)
Solutions of freshwater fish and amphibians to maintain osmotic balance
do not drink water, kidney produces large amounts of dilute urine, salt is actively pumped through skin by chloride cells (trying to maintain levels of salt)
Stenohaline (narrow salt)
cannot tolerate a large range of salinity and live in constant environments (marine or freshwater)
Euryhaline (wide salt)
Can tolerate a large range of salinity and can move between marine and freshwater environments
Diadromous
migrate between fresh and salt water
catadromous
life mostly in freshwater than migrate to spawn in saltwater (Atlantic eel)
anadromous
most adult life in saltwater then migrate to spawn in freshwater (salmon)
Osmostic regulation of terrestrial vertebrates
1) produce concentrated urine 2) produce very dry feces 3) decrease evaporative water loss through lungs and skin
10-20% of their body mass
Mammals cannot tolerate losses of water over __________.